- Brown bear
Temporal range: Late Pleistocene – Recent
A Kodiak bear (U. arctos middendorffi) in Katmai National Park, Alaska Conservation status Scientific classification Kingdom: Animalia Phylum: Chordata Class: Mammalia Order: Carnivora Family: Ursidae Genus: Ursus Species: U. arctos Binomial name Ursus arctos
Ursus arctos range map.
The brown bear (Ursus arctos) is a large bear distributed across much of northern Eurasia and North America. It can weigh from 300 to 780 kilograms (660 to 1,700 lb) and its largest subspecies, the Kodiak Bear, rivals the polar bear as the largest member of the bear family and as the largest land-based predator.
There are several recognized subspecies within the brown bear species. In North America, two types are generally recognized, the coastal brown bear and the inland grizzly, and the two types could broadly define all brown bear subspecies. Grizzlies weigh as little as 350 lb (159 kg) in Yukon, while a brown bear, living on a steady, nutritious diet of spawning salmon, from coastal Alaska and Russia can weigh 1,500 lb (682 kg). The exact number of overall brown subspecies remains in debate.
While the brown bear's range has shrunk, and it has faced local extinctions, it remains listed as a least concern species by the IUCN, with a total population of approximately 200,000. Its principal range countries are Russia, the United States (mostly in Alaska), Canada, the Carpathian region (especially Romania, but also Ukraine, Slovakia, and so on), the Balkans, Sweden and Finland, where it is the national animal. The brown bear is the most widely distributed of all bears.
- 1 Naming and etymology
- 2 Taxonomy and evolution
- 3 Physical description
- 4 Distribution and habitat
- 5 Behavior
- 6 Relationship with humans
- 7 See also
- 8 References
- 9 External links
Naming and etymology
The brown bear is sometimes referred to as the bruin, from Middle English, based on the name of the bear in History of Reynard the Fox, translated by William Caxton, from Middle Dutch bruun or bruyn, meaning brown (the color). During the Old West, the grizzly was termed "Old Ephraim" and sometimes as "Moccasin Joe".
Taxonomy and evolution
Brown bears are thought to have evolved from Ursus etruscus. The oldest fossils occur in China from about 0.5 million years ago. They entered Europe about 250,000 years ago, and North Africa shortly after. Brown bear remains from the Pleistocene period are common in the British Isles, where it is thought they outcompeted cave bears. The species entered Alaska 100,000 years ago, though they did not move south until 13,000 years ago. It is thought brown bears were unable to migrate south until the extinction of the much larger Arctodus simus. Several paleontologists suggest the possibility of two separate brown bear migrations: grizzlies are thought to stem from narrow-skulled bears which migrated from northern Siberia to central Alaska and the rest of the continent, while Kodiak bears descend from broad-skulled bears from Kamchatka, which colonized the Alaskan peninsula. Brown bear fossils discovered in Ontario, Ohio, Kentucky and Labrador show the species occurred farther east than indicated in historic records.
There is little agreement on classification of brown bears. Some systems have proposed as many as 90 subspecies, while recent DNA analysis has identified as few as five clades. DNA analysis recently revealed that the identified subspecies of brown bears, both Eurasian and North American, are genetically quite homogeneous, and that their genetic phylogeography does not correspond to their traditional taxonomy. As of 2005, 16 subspecies have been recognized. The subspecies have been listed as follows:
Subspecies name Image Distribution Description Ursus arctos arctos – Eurasian brown bear Europe, Caucasus, Siberia (except the east) and Mongolia A predominantly dark colored (rarely light colored), moderately-sized subspecies with dark claws, the Eurasian browns occurring in Siberia are larger than their European counterparts, as they are hunted less. Where found in Europe, primarily a forest creature Ursus arctos alascensis Alaska Ursus arctos beringianus – Kamchatka brown bear (or Far Eastern brown bear) Kamchatka Peninsula and Paramushir Island This is a very large, dark colored form. Light colored forms are encountered less than in European-Siberian subspecies. The claws are dark; it is thought to be the ancestor of U. a. middendorffi. Ursus arctos californicus – California golden bear (extinct) Ursus arctos collaris – East Siberian brown bear East Siberia from the Yenisei River to the Altai Mountains, also occurs in northern Mongolia A predominantly dark form, it is intermediate in size between U. a. arctos and U. a. beringianus, with a proportionately larger skull. Ursus arctos crowtheri – Atlas bear (extinct) Ursus arctos dalli Ursus arctos horribilis – Grizzly bear Western Canada, Alaska, and the northwestern United States, historically existed in Great Plains Grizzlies are identified by a medium to dark brown coat with gray, or "grizzled" tips on the fur. Smaller than the coastal bear, a grizzly typically weighs up to 800 lb (364 kg) in inland areas, with bears in the Yukon region weighing as little as 350 lb (159 kg). Coastal bears may be nearly twice a mountain grizzly's weight. Highly adaptable: can live in montane pine forests, temperate rainforest, arid scrubland, and prairie. Ursus arctos isabellinus – Himalayan brown bear Nepal, Pakistan, and Northern India Having a reddish-brown or sandy coat color, this bear is smaller than most other brown bears found on the Asian continent. Ursus arctos lasiotus – Ussuri brown bear (or Amur brown bear, black grizzly or horse bear) Russia: Southern Kuril Islands, Sakhalin, Maritime territory, and the Ussuri/Amur river region south of the Stanovoy Range, China: northeastern Heilongjiang, Japan: Hokkaidō This bear is thought to be the ancestor of U. a. horribilis. Ursus arctos middendorffi – Kodiak bear Kodiak, Afognak, Shuyak Islands (Alaska) This is the largest subspecies of brown bear, with other coastal brown bears reaching as big. Ursus arctos nelsoni – Mexican grizzly bear (extinct) Northern Mexico, including Chihuahua, Coahuila, and Sonora, southwestern United States including southern ranges of Texas, Arizona, and New Mexico This bear is believed extinct due to cattle ranching in both the United States and Mexico. Distinct in its ability to survive arid conditions, it could live in both montane pine forests of Mexico and canyonlands of Sonoran Desert. Ursus arctos pruinosus – Tibetan blue bear Western China and Tibet This is a moderately-sized subspecies with long and shaggy fur. Both dark and light variants are encountered, with intermediate colors predominating. The fur around the neck is light, and forms a "collar". The skull is distinguished its relatively flattened choanae, an arch-like curve of the molar row and large teeth. Ursus arctos sitkensis Baranof Island Appearing to be more closely related to the polar bear than to other brown bears, this species is called "clade I" by Waits, and others., and is part of the subspecies identified as U. a. sitkensis, by Hall and as U. a. dalli by Kurtén. Ursus arctos stikeenensis Ursus arctos syriacus – Syrian brown bear Occurs in the trans-Caucasus, Syria, Iraq,Turkey (Asia Minor), Iran, Afghanistan, western Himalayas and the Pamir-Alai and Tien Shan mountains, probable historical presence in Israel The Syrian is a light colored, moderate to small-sized subspecies with light claws. Ursus arctos marsicanus – Marsican brown bear (not officially recognised) Marsica (a small zone in Central Italy)
HybridsMain article: Grizzly–polar bear hybrid
A grizzly–polar bear hybrid (known as a pizzly Bear or grolar bear) is a rare ursid hybrid resulting from a union of a brown bear and a polar bear. It has occurred both in captivity and in the wild. In 2006, the occurrence of this hybrid in nature was confirmed by testing the DNA of a strange-looking bear that had been shot in the Canadian arctic. Previously, the hybrid had been produced in zoos, and was considered a "cryptid" (a hypothesized animal for which there is no scientific proof of existence in the wild).
Formerly considered subspecies
Image Distribution Description Ursus arctos pyrenaicus – Iberian brown bear, sometimes called Cantabrian brown bear, now considered Ursus arctos arctos - European brown bear See photographs in Eroski article (in Spanish, also available in Catalan, Basque and Galician) and in Fauna Ibérica. Oso pardo ibérico (Ursus arctos pyrenaicus), in Spanish Iberian Peninsula, primarily the Cantabrian Mountains and hills in Galicia, and the Pyrenees Until recently, this bear was considered a separate subspecies. Today, it is considered to belong to the U. arctos arctos subspecies. Scientific evidence based on DNA studies would furthermore indicate the European brown bear can be divided into two distinct lineages. "There is a clear division into two main mitochondrial lineages in modern European brown bear populations. These populations are divided into those carrying an eastern lineage (clade IIIa, Leonard et al. 2000), which is composed of Russian, northern Scandinavian and eastern European populations, and those carrying a western lineage (clade I, Leonard et al. 2000), which is composed of two subgroups, one believed to originate from the Iberian Peninsula, including southern Scandinavian bears and the Pyreneean populations; and the other from the Italian–Balkan peninsulas (Taberlet et al. 1994; see however Kohn et al. 1995). In addition, based on the subfossil record in northwestern Moldova and mitochondrial DNA data from modern populations, a Carpathian refuge has also been proposed (Sommer & Benecke 2005; Saarma et al. 2007)."
The brown bear is the largest animal on the Iberian Peninsula, although one of the smallest of the brown bears, since males rarely surpass 180 kg and adult females average 130 kg. Their fur varies from a pale cream color to dark brown, but always with a distinctively darker, nearly black tone at the paws and a yellowish tinge at the tip of each hair. The brown bear population is considered endangered in Spain.
The brown bear population in the Pyrenees stems mostly from bears reintroduced from Slovenia, with one or two remaining original males.
For further information, see Fundación Oso Pardo, with distribution maps and population from 2008 (in Spanish) and DEPANA, with detailed distribution maps and census information from 2009 (in Catalan). See also Staying out in the cold: glacial refugia and mitochondrial DNA phylogeography in ancient European brown bears, "Molecular Ecology", 2007, and The Cantabrian Brown Bear Ursus arctos arctos at the cAnfab site (in English).
Brown bears have very large and curved claws, those present on the forelimbs being longer than those on the hind limbs. They may reach 5 to 6 centimetres (2.0 to 2.4 in) and sometimes 7 to 10 centimetres (2.8 to 3.9 in) along the curve. They are generally dark with a light tip, with some forms having completely light claws. Brown bear claws are longer and straighter than those of American black bears. The claws are blunt, while those of a black bear are sharp.
Adults have massive, heavily built concave skulls, which are large in proportion to the body. The forehead is high and rises steeply. The projections of the skull are well developed when compared to those of Asian black bears: the latter have sagittal crests not exceeding more than 19–20% of the total length of the skull, while the former have sagittal crests comprising up to 40–41% of the skull's length. Skull projections are more weakly developed in females than in males. The braincase is relatively small and elongated. There is a great deal of geographical variation in the skull, and presents itself chiefly in dimensions. Grizzlies, for example, tend to have flatter profiles than European and coastal American brown bears. Skull lengths of Russian bears tend to be 31.5 to 45.5 centimetres (12.4 to 17.9 in) for males, and 27.5 to 39.7 centimetres (10.8 to 15.6 in) for females. The width of the zygomatic arches in males is 17.5 to 27.7 centimetres (6.9 to 11 in), and 14.7 to 24.7 centimetres (5.8 to 9.7 in) in females. Brown bears have very strong teeth: the incisors are relatively big and the canine teeth are large, the lower ones being strongly curved. The first three molars of the upper jaw are underdeveloped and single crowned with one root. The second upper molar is smaller than the others, and is usually absent in adults. It is usually lost at an early age, leaving no trace of the alveolus in the jaw. The first three molars of the lower jaw are very weak, and are often lost at an early age. Although they have powerful jaws, brown bear jaws are incapable of breaking large bones with the ease of spotted hyenas.
The dimensions of brown bears fluctuate very greatly according to sex, age, individual, geographic location, and season. The normal range of physical dimensions for a brown bear is a head-and-body length of 1.7 to 2.8 m (5.6 to 9.2 ft) and a shoulder height of 70 to 150 cm (28 to 59 in). The tail is relatively short, ranging from 6 to 22 cm (2.4 to 8.7 in) in length. The smallest subspecies is the Eurasian brown bear (U. a. arctos), whose mature females weigh as little as 90 kg (200 lb). Barely larger, grizzly bears (U. a. horribilis) from the Yukon region (which are a third smaller than most grizzlies) can weigh as little as 100 kg (220 lb) in the spring and the Syrian brown bear (U. a. syriacus), with mature females weighing as little as 150 kg (330 lb). The largest subspecies are the Kodiak bear (U. a. middendorffi) and the Kamchatka brown bear (U. a. beringianus), although bears from other coastal regions of eastern Asia and western North America can be comparably large. It is not unusual for large males in coastal regions to stand over 3.1 m (10 ft) tall while on their hind legs and to weigh up to 680 kg (1,500 lb). The heaviest recorded brown bear weighed over 1,150 kilograms (2,500 lb). Most brown bears fall somewhere between these extreme high and low weights, although even the relatively small Eurasian brown bear can weigh up to 481 kg (1,060 lb).
Brown bears have long, thick fur, with a moderately long mane at the back of the neck. In India, brown bears can be reddish with silver tips, while in China, brown bears are bicolored with a yellow-brown or whitish cape across the shoulders. North American grizzlies can be dark brown (almost black) to cream (almost white) or yellowish brown. Black hairs usually have white tips. The winter fur is very thick and long, especially in northern subspecies, and can reach 11 to 12 centimetres (4 to 5 in) at the withers. The winter hairs are thin, yet rough to the touch. The summer fur is much shorter and sparser, and its length and density varies geographically.
Distribution and habitat
There are about 200,000 brown bears in the world. The largest populations are in Russia with 120,000, the United States with 32,500, and Canada with 21,750. About 95% of the brown bear population in the United States is in Alaska, though in the lower 48 states, they are repopulating slowly but steadily along the Rockies and the western Great Plains. Although many people hold the belief some brown bears may be present in Mexico and the Atlas Mountains of Morocco, both are almost certainly extinct. The last Mexican brown bear was shot in 1960. In Europe, there are 14,000 brown bears in ten fragmented populations, from Spain (estimated at only 20-25 animals in the Pyrenees in 2010, in a range shared between France, Spain and Andorra, and some 85-90 animals in Asturias, Cantabria, Galicia and León, in the Picos de Europa and adjacent areas in 2003 and some 100 animals in 2005) in the west, to Russia in the east, and from Sweden and Finland in the north to Romania (4000–5000), Bulgaria (900–1200), Slovakia (with about 600–800 animals), Slovenia (500-700 animals) and Greece (with about 200 animals) in the south. They are extinct in the British Isles, extremely threatened in France and Spain, and in trouble over most of Central Europe. The Carpathian brown bear population of Romania is the largest in Europe outside Russia, estimated at 4,500 to 5,000 bears, although declining alarmingly due to overhunting. There is also a smaller brown bear population in the Carpathian Mountains in Ukraine (estimated at about 200 in 2005), Slovakia and Poland (estimated at about 100 in 2009 in the latter country). The total Carpathian population is estimated at about 8,000. Northern Europe is home to a large bear population, with an estimated 2,500 (range 2,350–2,900) in Sweden, about 1,600 in Finland, about 700 in Estonia and 70 in Norway. Another large and relatively stable population of brown bears in Europe, consisting of 2,500–3,000 individuals, is the Dinaric-Pindos (Balkans) population, with contiguous distribution in northeast Italy, Slovenia, Croatia, Bosnia and Herzegovina, Serbia, Montenegro, Macedonia, Albania, Bulgaria and Greece.
Brown bears were once native to Asia, the Atlas Mountains in Africa, Europe, and North America, but are now extinct in some areas, and their populations have greatly decreased in other areas. They prefer semiopen country, usually in mountainous areas.
Brown bears live in Alaska, east through the Yukon and Northwest Territories, south through British Columbia and through the western half of Alberta. Small populations exist in the Greater Yellowstone Ecosystem of northwest Wyoming (with about 600 animals), the Northern Continental Divide Ecosystem of northwest Montana (with about 750 animals), the Cabinet-Yaak Ecosystem of northwest Montana and northeast Idaho (with about 30–40 animals), the Selkirk Ecosystem of northeast Washington and northwest Idaho (with about 40–50 animals), and the North Cascades Ecosystem of northcentral Washington (with about 5–10 animals). These five ecosystems combine for a total of roughly 1,470 wild grizzlies still persisting in the contiguous United States. Unfortunately, these populations are isolated from each other, inhibiting any genetic flow between ecosystems. This poses one of the greatest threats to the future survival of the grizzly bear in the contiguous United States.
In Asia, brown bears are found in most of Russia, parts of the Middle East, and in a small area of Manchuria in China. They can also be found on the island of Hokkaidō in Japan, western China, and parts of North Korea, Afghanistan and India.
The population of brown bears in the Pyrenees mountain range between France and Spain is so low, estimated at 14 to 18, with a shortage of females, that bears, mostly female, from Slovenia were released in spring 2006 to reduce the imbalance and preserve the species' presence in the area, despite protests from French farmers.
A small population of brown bears (Ursus arctos marsicanus) still lives in central Italy (Apennine mountains, Abruzzo and Latium), with no more than 70 individuals, protected by strong laws, but endangered by the human presence in the area.
In Arctic areas, the potential habitat of the brown bear is increasing. The warming of that region has allowed the species to move farther north into what was once exclusively the domain of the polar bear. In non-Arctic areas, habitat loss is blamed as the leading cause of endangerment, followed by hunting.
North American brown bears, or grizzly bears, seem to prefer open landscapes, whereas in Eurasia they inhabit mostly dense forests. It is thought the Eurasian bears which colonized America were tundra-adapted. This is indicated by brown bears in the Chukotka Peninsula on the Asian side of Bering Strait, which are the only Asian brown bears to live year-round in lowland tundra like their North American cousins.
The brown bear is primarily nocturnal. In the summer, it gains up to 180 kilograms (400 lb) of fat, on which it relies to make it through winter, when it becomes very lethargic. Although they are not full hibernators, and can be woken easily; both sexes like to den in a protected spot, such as a cave, crevice, or hollow log, during the winter months. Brown bears are mostly solitary, although they may gather in large numbers at major food sources and form social hierarchies based on age and size. Adult male bears are particularly aggressive and are avoided by adolescent and subadult males. Female bears with cubs rival adult males in aggression, and are more intolerant of other bears than single females. Young adolescent males tend to be least aggressive, and have been observed in nonagonistic interactions with each other. Dominance between bears is asserted by making a frontal orientation, showing off canines, muzzle twisting and neck stretching to which a subordinate will respond with a lateral orientation, by turning away and dropping the head and by sitting or laying down. During combat, bears use their paws to strike their opponents in the chest or shoulders and bite the head or neck. In his Great Bear Almanac, Gary Brown lists 11 different sounds bears produce in 9 different contexts. Sounds expressing anger or aggravation include growls, roars, woofs, champs and smacks, while sounds expressing nervousness or pain include woofs, grunts and bawls. Sows will bleat or hum when communicating with their cubs.
The mating season is from late May to early July. Being serially monogamous, brown bears remain with the same mate from several days to a couple of weeks. Females mature sexually between the age of 5 and 7 years, while males usually mate a few years later, when they are large and strong enough to successfully compete with other males for mating rights.
Males, however, take no part in raising their cubs – parenting is left entirely to the females.
Through the process of delayed implantation, a female's fertilized egg divides and floats freely in the uterus for six months. During winter dormancy, the fetus attaches to the uterine wall. The cubs are born eight weeks later, while the mother sleeps. If the mother does not gain enough weight to survive through the winter, the embryo does not implant and is reabsorbed into the body. The average litter has one to four cubs, usually two. There have been cases of bears with five cubs, although females sometimes adopt stray cubs. Older females tend to give birth to larger litters. The size of a litter also depends on factors such as geographic location and food supply. At birth, the cubs are blind, toothless, hairless, and weigh less than 450 grams (1.0 lb). They feed on their mother's milk until spring or even early summer, depending on climate conditions. At this time, the cubs weigh 7 to 9 kilograms (15 to 20 lb) and have developed enough to follow her and begin to forage for solid food.
Cubs remain with their mother from two to four years, during which time they learn survival techniques, such as which foods have the highest nutritional values and where to obtain them; how to hunt, fish, and defend themselves; and where to den. The cubs learn by following and imitating their mother's actions during the period they are with her. Brown bears practice infanticide. An adult male bear may kill the cubs of another bear either to make the female sexually receptive or simply for consumption. Cubs flee up a tree when they see a strange male bear, and the mother defends them, even though the male may be twice her size.
They are omnivores and feed on a variety of plant products, including berries, roots, and sprouts, and fungi, as well as meat products such as fish, insects, and small mammals. Despite their reputation, most brown bears are not highly carnivorous, as they derive up to 90% of their dietary food energy from vegetable matter. Their jaw structure has evolved to fit their dietary habits. Their diet varies enormously throughout their differing areas based on opportunity. For example, bears in Yellowstone eat an enormous number of moths during the summer, sometimes as many as 40,000 in a day, and may derive up to half of their annual food energy from these insects. In the Kamchatka peninsula and parts of coastal Alaska, brown bears feed mostly on spawning salmon, whose nutrition and abundance explain the enormous size of the bears in these areas, though, these bears very rarely hunt large animals. Brown bears also occasionally prey on large mammals, such as deer (including elk, moose and caribou), bighorn sheep, mountain goats, bison and muskoxen. When brown bears attack these animals, they tend to choose the young ones, as they are easier to catch. When hunting, the bear pins its prey to the ground and then tears and eats it alive. On rare occasions, bears kill by hitting their prey with their powerful forearms, which can break the necks and backs of large prey, such as moose. They also feed on carrion, and use their size to intimidate other predators, such as wolves, cougars, tigers, and black bears from their kills.
Interspecific predatory relationships
Brown bears often use their large size for intimidation when a kill or a territory is in dispute with another large predator and they are normally dominant in such interactions. Sometimes, the conflict will escalate to the point of violence, but usually threat displays are sufficient, since most animals try to avoid potential bodily harm. However, the massive strength and size of the brown bear will usually result in it winning violent conflicts, even against wolf packs and Siberian Tigers. In situations where the interspecies conflict turns deadly, brown bears may also eat the competitor, despite it not being the primary reason for attack.
Brown bears regularly intimidate wolves away from their kills. In Yellowstone National Park, brown bears pirate wolf kills so often, Yellowstone's Wolf Project Director Doug Smith wrote, "It's not a matter of if the bears will come calling after a kill, but when." Though conflict over carcasses is common, on rare occasions the two predators tolerate each other on the same kill. Given the opportunity, both species prey on the other's cubs. Conclusively, the individual power of the bear against the collective strength of the wolf pack usually results in a long battle for kills or domination. In some areas, the brown bear also regularly displaces cougars from their kills.
Adult bears are generally immune from predatory attacks from anything other than other bears. Some bears emerging from hibernation seek tigers to steal their kills. Indeed, Russian researchers have identified "satellite bears" who "follow tigers over extensive periods of time, sequentially usurping kills"; these bears were observed tracking tigers in spring snow and regularly usurped their kills. In the Russian Far East, brown bears along with smaller Asiatic black bears constitute 5–8% of the diet of Siberian tigers. In particular, the brown bear's input is estimated to be 1.0–1.5% in one source. However, another source states that such attacks are rare and do not have any actual significance because Siberian tigers are almost extinct. Siberian tigers most typically attack brown bears in the winter in the hibernaculum or in the late autumn and early spring, and when ungulate populations decrease. Adult bears, generally smaller ones, are sometimes vulnerable to tiger attacks and have been killed in their dens in winter, with the tiger taking advantage of the bear's hibernating condition. There are also records of bears killing tigers, including fully grown adult males and tigers whose sex and age were not specified, either in self-defense, or in disputes over kills or for consumption.
Brown bears usually dominate other bear species in areas where they coexist. Due to their smaller size, American black bears are at a competitive disadvantage to brown bears in open, unforested areas. Although displacement of black bears by brown bears has been documented, actual interspecific killing of black bears by brown bears has only occasionally been reported. Confrontation is mostly avoided due to the black bear's diurnal habits and preference for heavily forested areas, as opposed to the brown bear's largely nocturnal habits and preference for open spaces. Brown bears may attack Asian black bears. They will eat the fruit dropped by the latter species from trees, as they themselves are too large and cumbersome to climb.
There has been a recent increase in interactions between brown bears and polar bears, theorized to be caused by climate change. Brown bears have been seen moving increasingly northward into territories formerly claimed by polar bears. Brown bears tend to dominate polar bears in disputes over carcasses, and dead polar bear cubs have been found in brown bear dens.
Relationship with humans
Bears become attracted to human-created food sources, such as garbage dumps, litter bins, and dumpsters; they venture into human dwellings or barns in search of food as humans encroach into bear habitats. In the U.S., bears sometimes kill and eat farm animals. When bears come to associate human activity with a "food reward", they are likely to continue to become emboldened; the likelihood of human-bear encounters increases, as they may return to the same location despite relocation. The saying, "a fed bear is a dead bear", has come into use to popularize the idea that allowing bears to scavenge human garbage, such as trash cans and campers' backpacks, pet food, or other food sources that draw the bear into contact with humans, can result in a bear's death.
Relocation of the bear has been used to separate the bear from the human environment, but it does not address the problem of the bear's newly-learned association of humans with food or the environmental situations which created the human-habituated bear. "Placing a bear in habitat used by other bears may lead to competition and social conflict, and result in the injury or death of the less dominant bear."
Yellowstone National Park, an enormous reserve located in the western United States, contains prime habitat for the grizzly bear (Ursus arctos horribilis), and due to the enormous number of visitors, human-bear encounters are common. The scenic beauty of the area has led to an influx of people moving into the area. In addition, because there are so many bear relocations to the same remote areas of Yellowstone, and because male bears tend to dominate the center of the relocation zone, female bears tend to be pushed to the boundaries of the region and beyond. As a result, a large proportion of repeat offenders, bears that are killed for public safety, are females. This creates a further depressive effect on an already endangered species. The grizzly bear is officially described as "threatened" in the U.S. Though the problem is most significant with regard to grizzlies, these issues affect the other types of brown bears as well.
In Europe, part of the problem lies with shepherds; over the past two centuries, many sheep and goat herders have gradually abandoned the more traditional practice of using dogs to guard flocks, which have concurrently grown larger. Typically, they allow the herds to graze freely over sizeable tracts of land. As bears reclaim parts of their range, they may eat livestock. In some cases, the shepherds shoot the bear, thinking their livelihood is under threat. Many are now better informed about the ample compensation available, and will make a claim when they lose livestock to a bear.
- The grizzly bear, sometimes called the silvertip bear, is listed as threatened in the contiguous United States. It is slowly repopulating in areas where it was previously extirpated, though it is still vulnerable.
- The California golden bear (Ursus arctos californicus) disappeared from the state of California in 1922, when the last one was shot in Tulare County. It is the official state animal.
- The Mexican grizzly bear is listed as an endangered species, but it may be extinct.
- In Canada, it is listed as vulnerable in Alberta, British Columbia, Northwest Territories, and Yukon Territory. Prairie populations of grizzly bear are listed as extirpated in Alberta, Manitoba, and Saskatchewan.
- The brown bear is a European Protected Species, given protection throughout the European Union.
- The brown bear is also the national animal of Finland.
- The brown bear is depicted on the reverse of the Croatian 5 kuna coin, minted since 1993.
There are an average of two fatal attacks by bears per year in North America. In Scandinavia, there are only four known cases since 1902 of bear encounters which have resulted in death. The two most common causes for bear attack are surprise and curiosity. Some types of bears, such as polar bears, are more likely to attack humans when searching for food, while American black bears are much less likely to attack.
The Alaska Science Center ranks the following as the most likely reasons for bear attacks:
- Invaded personal space (this includes a mother bear protecting her young)
- Predatory intent
- Hunting wounded
- Carcass defense
- Provoked charge
Aggressive behavior in brown bears is favored by numerous selection variables. Unlike the smaller black bears, adult brown bears are too large to escape danger by climbing trees, so they respond to danger by standing their ground and warding off their attackers. Increased aggressiveness also assists female brown bears in better ensuring the survival of their young to reproductive age. Mothers defending cubs are the most prone to attacking, being responsible for 70% of brown bear-caused human fatalities in North America.
Attacks on humans
As a rule, brown bears seldom attack humans on sight, and usually avoid people. They are, however, unpredictable in temperament, and will attack if they are surprised or feel threatened. Sows with cubs account for the majority of injuries and fatalities in North America. Habituated or food-conditioned bears can also be dangerous, as their long-term exposure to humans causes them to lose their natural shyness, and, in some cases, to associate humans with food. Small parties of one or two people are more often attacked than large groups, with only one known case of an attack on a group of six or more. In that instance, it is thought that due to surprise the bear may not have recognized the size of the group. In contrast to injuries caused by American black bears, which are usually minor, brown bear attacks tend to result in serious injury and, in some cases, death. In the majority of attacks resulting in injury, brown bears precede the attack with a growl or huffing sound, and seem to confront humans as they would when fighting other bears: they rise up on their hind legs, and attempt to "disarm" their victims by biting and holding on to the lower jaw to avoid being bitten in turn. Due to the bears' enormous physical strength, even a single bite or swipe can be deadly, as in tigers, with some human victims having had their heads completely crushed by a bear bite. Most attacks occur in the months of July, August and September, the time when the number of outdoor recreationalists, such as hikers or hunters, is higher. People who assert their presence through noises tend to be less vulnerable, as they alert bears to their presence. In direct confrontations, people who run are statistically more likely to be attacked than those who stand their ground. Violent encounters with brown bears usually last only a few minutes, though they can be prolonged if the victims fight back.
Attacks on humans are considered extremely rare in the former Soviet Union, though exceptions exist in districts where they are not pursued by hunters. Siberian bears, for example, tend to be much bolder toward humans than their shyer, more persecuted European counterparts. In 2008, a platinum mining compound in the Olyotorsky district of northern Kamchatka was besieged by a group of 30 bears, who killed two guards and prevented workers from leaving their homes. Ten people a year are killed by brown bears in Russia. In Scandinavia, only three fatal attacks were recorded in the 20th century.
In Japan, a large brown bear nicknamed "Kesagake" (袈裟懸け, "kesa-style slasher") made history for causing the worst bear attack in Japanese history at Tomamae, Hokkaidō during numerous encounters in December, 1915. It killed seven people (including one pregnant woman) and wounded three others (with possibly another three previous fatalities to its credit) before being gunned down after a large-scale beast-hunt. Today, there is still a shrine at Rokusensawa (六線沢), where the event took place, in memory of the victims of the incident.
Native American tribes sympatric to brown bears often viewed them with a mixture of awe and fear. North American brown bears were so feared by the Natives, they were rarely hunted, especially alone. When Natives hunted grizzlies, the act was done with the same preparation and ceremoniality as intertribal warfare, and was never done except with a company of 4–10 warriors. The tribe members who dealt the killing blow were highly esteemed among their compatriots. Californian Indians actively avoided prime bear habitat, and would not allow their young men to hunt alone, for fear of bear attacks. During the Spanish colonial period, some tribes, instead of hunting grizzlies themselves, would seek aid from European colonists to deal with problem bears. Many authors in the American west wrote of Natives or voyagers with lacerated faces and missing noses or eyes due to attacks from grizzlies. Within Yellowstone National Park, injuries caused by grizzly attacks in developed areas averaged approximately one per year during the 1930s through to the 1950s, though it increased to four per year during the 1960s. They then decreased to one injury every two years during the 1970s. Between 1980 and 2002, there have been only two human injuries caused by grizzly bears in a developed area. Though grizzly attacks were rare in the backcountry before 1970, the number of attacks increased to an average of approximately one per year during the 1970s, 1980s, and 1990s.
History of defense from bears
A study by Canadian and US researchers has found pepper spray to be more effective at stopping aggressive bear behavior than guns, working in 92% of studied incidents versus 67% for guns. Carrying pepper spray is highly recommended by many authorities when traveling in bear country; however, carrying two means of deterrent, one of which is a large caliber gun, is also advised. Solid shotgun slugs, or three buckshot rounds, or a semiautomatic pistol of .45 caliber or more is suggested if a heavy hunting rifle is not available. Guns remain a viable, last resort option to be used in defense of life from aggressive bears. Too often, people do not carry a proper caliber weapon to neutralize the bear. According to the Alaska Science Center, a 12 gauge shotgun with slugs has been the most effective weapon. There have been fewer injuries as a result of only carrying lethal loads in the shotgun, as opposed to deterrent rounds. State of Alaska Defense of Life or Property (DLP) laws require one to report the kill to authorities, and salvage the hide, skull, and claws.
Campers are often told to wear bright colored red ribbons and bells, and carry whistles to ward off bears. They are told to look for grizzly scat in camping areas, and be careful to carry the bells and whistles in those areas. Grizzly scat is difficult to differentiate from black bear scat, as diet is in a constant state of flux depending on the availability of seasonal food items. If a bear is killed near camp, the bear's carcass must be adequately disposed of, including entrails and blood, if possible. Failure to move the carcass has often resulted in it attracting other bears and further exacerbating a bad situation. Moving camps immediately is another recommended method.
State of Alaska Department of Natural Resources website states, "Select a gun that will stop a bear (12-gauge shotgun or .300 mag rifle)."
Many Native American tribes both respected and feared the brown bear, even thinking of it as a god. One tale tells of how the black bear was a creation of the Great Spirit, while the grizzly was created by the Evil Spirit. In Kwakiutl mythology, black and brown bears became enemies when Grizzly Bear Woman killed Black Bear Woman for being lazy. Black Bear Woman's children, in turn, killed Grizzly Bear Woman's own cubs.
The grizzly bear is the state animal of both Montana and California.
Brown bears often figure into the literature of Europe and North America, in particular that which is written for children. "The Brown Bear of Norway" is a Scottish fairy tale telling the adventures of a girl who married a prince magically turned into a bear, and who managed to get him back into a human form by the force of her love and after many trials and difficulties. With "Goldilocks and the Three Bears", a story from England, the three bears are usually depicted as brown bears. In German speaking countries, children are often told the fairytale of Snow White and Rose Red; the handsome prince in this tale has been transfigured into a brown bear. In the United States, parents often read their preschool age children the book Brown Bear, Brown Bear, What Do You See? to teach them their colors and how they are associated with different animals.
The Russian bear is a common national personification for Russia (as well as the Soviet Union). The brown bear is also Germany's and Finland's national animal.
The school mascot for Brown University, the University of California, Los Angeles, the University of California, Berkeley, and the University of Alberta is the brown bear.
The coat of arms of Madrid depicts a bear reaching up into a madroño or strawberry tree (Arbutus unedo) to eat some of its fruit, whereas the Swiss city of Bern's coat of arms also depicts a bear and the city's name is popularly thought to derive from the German word for bear.
In the town of Prats de Molló, in Vallespir, Northern Catalonia, a "bear festival" (festa de l'ós) is celebrated annually at the beginning of spring, in which the locals dress up as bears, cover themselves with soot or coal and oil, and "attack" the onlookers, attempting to get everyone dirty. The festival ends with the ball de l'os (bear dance).
- European brown bear
- Cantabrian brown bear
- Environmental Centre ARCTUROS
- List of fatal bear attacks in North America
- Sankebetsu brown bear incident
- Kodiak bear
- Grizzly bear
- ^ McLellan, B.N., Servheen, C. & Huber, D. (2008). Ursus arctos. In: IUCN 2008. IUCN Red List of Threatened Species. Downloaded on 27 January 2009.
- ^ "Polar bear, (Ursus maritimus)" (PDF). U.S. Fish and Wildlife service. http://www.fws.gov/endangered/esa-library/pdf/polar_bear.pdf. Retrieved 2008-03-22. "Appearance. The polar bear is the largest member of the bear family, with the exception of Alaska's Kodiak brown bears, which equal polar bears in size." (Overview page)
- ^ Craig Boddington, illustrations by Ken Carlson (2004). Fair Chase in North America (illustrated ed.). Boone and Crockett Club. p. 45. ISBN 0940864479, 9780940864474. http://books.google.com/?id=Rri8xdwgdQ0C&pg=PA45&dq=largest+predator+bear.
- ^ American Heritage Dictionary of the English Language: Fourth Edition. 2000.
- ^ Hunting the Grisly and other Sketches. Fullbooks.com. Retrieved on 2011-09-15.
- ^ a b c d Bruce McLellan and David C. Reiner (1994). A Review of bear evolution Int. Conf. Bear Res. and Manage. 9(1):85–96
- ^ a b c Valerius Geist Did Large Predators keep Humans out of North America?. (PDF) . Retrieved on 2011-09-15.
- ^ U.S. Fish and Wildlife Service (2006-11-17). "Endangered and Threatened Wildlife and Plants; Designating the Greater Yellowstone Ecosystem Population of Grizzly Bears as a Distinct Population Segment; Removing the Yellowstone Distinct Population Segment of Grizzly Bears From the Federal List of Endangered and Threatened Wildlife" (PDF). Federal Register / Vol. 70, No. 221. pp. 69854–69884. Archived from the original on August 25, 2006. http://web.archive.org/web/20060825042929/http://www.bearinfo.org/YESFedRegister.pdf. Retrieved August 1, 2006.
- ^ a b Lisette P. Waits, Sandra L. Talbot, R.H. Ward and G. F. Shields (April 1998). "Mitochondrial DNA Phylogeography of the North American Brown Bear and Implications for Conservation". Conservation Biology. pp. 408–417. http://www.cnrhome.uidaho.edu/documents/Waits%20et%20al%201998%20cb.pdf&pid=78496&doc=1. Retrieved August 1, 2006.
- ^ Wozencraft, W. Christopher (16 November 2005). "Order Carnivora (pp. 532-628)". In Wilson, Don E., and Reeder, DeeAnn M., eds. Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Baltimore: Johns Hopkins University Press, 2 vols. (2142 pp.). ISBN 978-0-8018-8221-0. OCLC 62265494. http://www.bucknell.edu/msw3/browse.asp?id=14000970.
- ^ a b c d e f g h i j k l m n o p q Mammals of the Soviet Union Vol.II Part 1a, SIRENIA AND CARNIVORA (Sea cows; Wolves and Bears), V.G Heptner and N.P Naumov editors, Science Publishers, Inc. USA. 1998. ISBN 1-886106-81-9
- ^ Ursus arctos alascensis Merriam, 1896 .Taxonomic Serial No.: 726985. Integrated Taxonomic Information System (ITIS). http://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=726985&print_version=PRT&source=to_print. . Also Wilson and Reeders, Mammal Species of the World, 3rd edition 
- ^ "Wild find: Half grizzly, half polar bear: Hunter bags what expert 'never thought would happen' in wild". MSNBC.MSN.com. May 11, 2006. http://msnbc.msn.com/id/12738644/?GT1=8199. Retrieved 2006-05-14.
- ^ James Mallet (2008). "Hybridization, ecological races and the nature of species: empirical evidence for the ease of speciation". Phil. Trans. R. Soc. B 363 (1506): 2971–2986. doi:10.1098/rstb.2008.0081. PMC 2607318. PMID 18579473. http://www.ucl.ac.uk/taxome/jim/pap/Mallet08%20Phil%20Trans.pdf.
- ^ Anthony D. Barnosky (2009). Island Press. ed. Heatstroke: Nature in an Age of Global Warming. Washington, DC: Island Press/Shearwater Books. ISBN 1597261971. http://books.google.com/?id=Di6SVZZjLAgC&pg=PA10&dq=Pizzly+Bear.
- ^ a b c "La recuperació del rei dels boscos" (in Basque, Catalan, Galician, Spanish). http://revista.consumer.es/web/ca/20050901/medioambiente/69893.php. Retrieved 2010-11-19.
- ^ Staying out in the cold: glacial refugia and mitochondrial DNA phylogeography in ancient European brown bears, "Molecular Ecology", Blackwell Publishing Ltd., 2007, p.2.
- ^ a b c d e Bear Anatomy and Physiology from Gary Brown's The Great Bear Almanac, Lyons & Burford, Publishers, 1993
- ^ a b ''The living animals of the world; a popular natural history with one thousand illustrations'' Volume 1: Mammals, by Cornish, C. J. (Charles John), 1858–1906; Selous, Frederick Courteney, 1851–1917; Johnston, Harry Hamilton, Sir, 1858–1927; Maxwell, Herbert, Sir, published by New York, Dodd, Mead and Company. Archive.org. Retrieved on 2011-09-15.
- ^ John M. Rensberger (1999). "Enamel microstructural specialization in the canine of the spotted hyena, Crocuta crocuta". Scanning Microscopy 13 (2–3): 343–361). http://ecmjournal.org/journal/smi/pdf/smi99-37.pdf.
- ^  (2011).
- ^ Wood, The Guinness Book of Animal Facts and Feats. Sterling Pub Co Inc (1983), ISBN 978-0-85112-235-9
- ^ The Bear Facts – Types of bears in the Yukon, Yukon Education Student Network
- ^ "Kodiak Bear Fact Sheet". Alaska Department of Fish and Game, Division of Wildlife Conservation. 2008. http://www.wildlife.alaska.gov/index.cfm?adfg=bears.trivia. Retrieved 2008-10-27.
- ^ Wood, Gerald (1983). The Guinness Book of Animal Facts and Feats. p. 256. ISBN 978-0-85112-235-9.
- ^ "Neixen quatre cries d'ós bru als Pirineus" (in Catalan; Spanish). http://www.elperiodico.cat/ca/noticias/societat/20100802/neixen-quatre-cries-dos-bru-als-pirineus/416750.shtml. Retrieved 2010-11-19.
- ^ "Situació preocupant de l'ós bru als Pirineus" (in Catalan). http://margalloecologistes.wordpress.com/2010/05/27/situacio-preocupant-de-los-bru-als-pirineus/. Retrieved 2010-11-19.
- ^ "La población del oso pardo en España se reduce a 90 ejemplares" (in Spanish). http://www.consumer.es/web/es/medio_ambiente/2003/06/26/62830.php. Retrieved 2010-11-19.
- ^ [dead link]
- ^ "Proact Local Campaigns: Carpathian Brown Bear". http://proaction.tripod.com/localcampaigns/id26.html. Retrieved 2010-11-19.
- ^ "Carpathian Brown Bear Management Plan [in Poland"]. http://carpathianbear.pl/en/projects/management-plan. Retrieved 2010-11-19.
- ^ "Brown Bear - Population & Distribution: A Truly International Species". http://wwf.panda.org/about_our_earth/species/profiles/mammals/brown_bear2/brownbear_population_distribution/. Retrieved 2010-11-19.
- ^ "Suurpedot.fi – Karhu" (in finnish). http://www.suurpedot.fi/www/fi/lajit/karhu/index.php. Retrieved 2010-10-22.
- ^ Bear Online Information System for Europe. Kora.ch. Retrieved on 2011-09-15.
- ^ "Ancient bear made early migration". BBC News. November 12, 2004. http://news.bbc.co.uk/2/hi/science/nature/4003325.stm. Retrieved January 2, 2010.
- ^ "Brown Bear Hunting in Russia". Russianbearhunt.com. http://www.russianbearhunt.com/. Retrieved 2009-10-10.
- ^ Egbert, Allan L.; Stokes, Allen W.; Egbert, A. L. (1974). "The social behaviour of brown bears on an Alaskan salmon stream" (PDF). International Conference Bear Res. and Manage 3: 41–56. JSTOR 3872753. http://www.bearbiology.com/fileadmin/tpl/Downloads/URSUS/Vol_3/Egbert_Stokes_Egbert_Vol_3.pdf.
- ^ DEREK STONOROV and ALLEN W. STOKES (1972) "Social Behavior of the Alaska Brown Bear" PANEL 4: BEAR BEHAVIOUR
- ^ Types of Bears – Information on Specific Bear Species (2009)
- ^ "Brown Bear Reproduction". Shadowofthebear.com. Archived from the original on May 1, 2008. http://web.archive.org/web/20080501092730/http://www.shadowofthebear.com/reproduction.html. Retrieved 2009-10-10.
- ^ Bellemain, Eva; Swenson, Jon E.; Taberlet, Pierre (2006). "Mating Strategies in Relation to Sexually Selected Infanticide in a Non-Social Carnivore: The Brown Bear". Ethology 112 (3): 238–246. doi:10.1111/j.1439-0310.2006.01152.x.
- ^ "Alaska Office of Economic Development". Dced.state.ak.us. http://www.dced.state.ak.us/oed/student_info/learn/bearwatching.htm. Retrieved 2009-10-10.
- ^ "Yellowstone Grizzly Bears Eat 40,000 Moths a Day In August". Yellowstonepark.com. http://www.yellowstonepark.com/MoreToKnow/ShowNewsDetails.aspx?newsid=163. Retrieved 2009-10-10.
- ^ French, S. P.; French, M. G. (1990). "Predatory behavior of grizzly bears feeding on elk calves in Yellowstone National Park, 1986–1988" (PDF). International Conf. Bear Res. and Manage 8: 335–341. http://www.bearbiology.com/fileadmin/tpl/Downloads/URSUS/Vol_8/French_French_8.pdf.
- ^ Betsy Downey. "Personal Encounter. Wolf-Grizzly interaction in Yellowstone National Park". International Wolf Center. http://www.wolf.org/wolves/news/iwmag/2006/spring/personalencounter.pdf.
- ^ Kerry M. Murphy, Gregory S. Felzien, Maurice G. Hornocker and Toni K. Ruth (1998). "Encounter Competition between Bears and Cougars: Some Ecological Implications". Ursus 10: 55–60. JSTOR 3873109.
- ^ Matthiessen, Peter; Hornocker, Maurice (2001). Tigers In The Snow. North Point Press. ISBN 0865475962.
- ^ a b Kerley, Linda; Goodrich, John, and Miquelle, Dale. "Bears and tigers in the Far East" International Bear News. 5 (2): p. 4.
- ^ Vratislav Mazak: Der Tiger. Nachdruck der 3. Auflage von 1983. Westarp Wissenschaften Hohenwarsleben, 2004 ISBN 3-89432-759-6
- ^ V.G. Heptner and N.P. Naumov. Mammals of the Soviet Union. Vol. 2 IIa 1998. New Delhi, India: Amerind Publishing. pp. 670-1
- ^ a b Mammals of the Soviet Union Volume II, by V.G Heptner & A.A. Sludskii p. 177
- ^ [dead link]
- ^ Seryodkin, I. V., J. M. Goodrich, A. V. Kostyrya, B. O. Schleyer, E. N. Smirnov, L. L. Kerley, and D. G. Miquelle. 2005. Relationship between tigers, brown bears, and Himalayan black bears. pp. 156–163 in D. G. Miquelle, E. N. Smirnov, and J. M. Goodrich (eds.), Tigers of Sikhote-Alin Zapovednik: Ecology and Conservation. Vladivostok, Russia: PSP.
- ^ V.G Heptner & A.A. Sludskii (1992). Mammals of the Soviet Union, Volume II, Part 2. Leiden u.a.: Brill. ISBN 9004088768.
- ^ Kerry A. Gunther, Mark J. Biel, Neil Anderson & Lisette Watts (2002). "Probable grizzly bear predation on an American black bear in Yellowstone National Park" (PDF). Ursus 13: 372–374. http://www.nps.gov/yell/naturescience/upload/predation.pdf.
- ^ Ivan V. Seryodkin et al. (2003). "Denning ecology of brown bears and Asiatic black bears in the Russian Far East". Ursus 14 (2): 153–161. JSTOR 3873015. http://www.bearbiology.com/fileadmin/tpl/Downloads/URSUS/Vol_14_2/Seryodkin_14_2.pdf.
- ^ The Intellectual observer: review of natural history, microscopic research, and recreative science, published by Groombridge., 1865
- ^ Dough O'Hara Polar bears, grizzlies increasingly gather on North Slope. Anchorage Daily News. April 24, 2005
- ^ "ABC News: Grizzlies Encroaching on Polar Bear Country". Abcnews.go.com. http://abcnews.go.com/Technology/DyeHard/Story?id=582243&page=3. Retrieved 2009-10-10.
- ^ "Relocation". Get Bear Smart Society. http://web.archive.org/web/20060513075036/http://www.bearsmart.com/managingBears/Relocation.html.
- ^ Wilson, Don E., and DeeAnn M. Reeder, eds.. "Ursus arctos californicus". Johns Hopkins University Press. http://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=726987. Retrieved April 1, 2008.
- ^ "History and Culture - State Symbols". California State Library. http://www.library.ca.gov/history/symbols.html. Retrieved 23 September 2011.
- ^ Bear Attacks: Their Causes and Avoidance, Stephen Herrero, revised edition, 2002.
- ^ a b WTom S. Smith, Ph.D. and Steven Herrero, Ph.D.. "Ursus arctos californicus". Alaska Science Center – Biological Science Office. http://www.absc.usgs.gov/research/brownbears/attacks/bear-human_conflicts.htm. Retrieved April 12, 2008.
- ^ Why are grizzly bears more aggressive than our black bears?. Digital Collegian. April 27, 2004
- ^ Lynn L. Rogers How Dangerous are Black Bears
- ^ a b c d Bear Attacks: Their Causes and Avoidance by Stephen Herrero, Hurtig Publishers Ltd./ Edmonton 1985
- ^ Alaska bear attack: NOLS kids did a 'phenomenal job', Craig Medred, Alaska Dispatch, 26 July 2011
- ^ WOLVES, BEARS AND HUMAN ANTI-PREDATOR ADAPTATIONS[dead link]
- ^ Bears besiege Russian mine after killing guards. The Times. July 24, 2008
- ^ Brown Bears of Russia. Dinets.travel.ru. Retrieved on 2011-09-15.
- ^ "Brown Bear". Tooth & Claw. http://www.toothandclaw.org.uk/species.asp. Retrieved 2008-01-05.
- ^ Bear Caused Human Injuries and Deaths In Yellowstone National Park. Yellowstone-bearman.com (2000-01-01). Retrieved on 2011-09-15.
- ^ Smith, Herrero, DeBruyn, Wilde (2008). "Spray more effective than guns against bears: study". North American Bear Center. http://www.bear.org/website/index.php?option=com_content&task=view&id=202&Itemid=123. Retrieved 2009-03-28.
- ^ Smith, Tom S.. "Brown Bear Projects at the Alaska Science Center". Alaska Science Center – Biological Science Office. http://www.absc.usgs.gov/research/brownbears/safety/safeconduct.htm. Retrieved 2008-06-02.
- ^ "Alaska State Troopers Press Release of Monday, November 19, 2007" (Case Number: 07-96958). Alaska Department of Public Safety. 2007-11-19. Archived from the original on April 17, 2008. http://web.archive.org/web/20080417183426/http://www.dps.state.ak.us/pio/dispatch/Trooper+Dispatches+of+11-19-2007.20071119.txt. Retrieved 2008-06-02.
- ^ Native American Animal Symbols – The Bear. Support-native-american-art.com (2004-01-22). Retrieved on 2011-09-15.
- ^ Folklore and Legends of the North American Indian compiled by Joshua B Lippincott, published by Abela Publishing Ltd, 2009, ISBN 0-9560584-6-9
- ^ Kwakiutl String Figures Julia P. Averkieva and Mark A. Sherman, UBC Press, 1992, ISBN 0-7748-0432-7
- Brown Bear profile from National Geographic
- IFAW Rebuilding the European Brown Bear Population
- Bear Hunting Altered Genetics More Than Ice Age Isolation
- Ancient Fossil Offers New Clues To Brown Bears Past
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