Cycas

Cycas
Leaves and male cone of Cycas revoluta
Scientific classification
Kingdom:	Plantae
Division:	Cycadophyta
Class:	Cycadopsida
Order:	Cycadales
Family:	Cycadaceae Persoon
Genus:	Cycas L.
Species
See Species Section

Cycas is the type genus and the only genus currently recognised in the cycad family Cycadaceae. About 95 species are currently accepted. The best-known species is Cycas revoluta, widely cultivated under the name "Sago Palm" or "King Sago Palm" due to its palm-like appearance although it is not a true palm. The generic name comes from Greek Koikas, and means "a kind of palm".

The genus is native to the Old World, with the species concentrated around the equatorial regions. It is native to eastern and southeastern Asia including the Philippines with 10 species (9 of which are endemic), eastern Africa (including Madagascar), northern Australia, Polynesia, and Micronesia. Australia has 26 species, while the Indo-Chinese area has about 30. The northernmost species (C. revoluta) is found at 31°N in southern Japan. The southernmost (C. megacarpa) is found at 26°S in southeast Queensland, Australia.

The plants are dioecious, and the family Cycadaceae is unique among the cycads in not forming seed cones on female plants, but rather a group of leaf-like structures each with seeds on the lower margins, and pollen cones on male individuals.

Bark of Cycas rumphii

The caudex is cylindrical, surrounded by the persistent petiole base. Most species form distinct branched or unbranched trunks but in some species the main trunk can be subterranean with the leaf crown appearing to arise directly from the ground. The leaves are pinnate (or more rarely bipinnate) and arranged spirally, with thick and hard keratinose. The leaflets are articulated, have midrib but lack secondary veins. Megasporophylls are not gathered in cones.

Often considered a living fossil, the earliest fossils of the genus Cycas appear in the Cenozoic although Cycas-like fossils that may belong to Cycadaceae extend well into the Mesozoic. Cycas is not closely related to other genera of cycads, and phylogenetic studies have shown that Cycadaceae is the sister-group to all other extant cycads.

The plant takes several years to grow, sexual reproduction takes place after 10 years of exclusive vegetative growth.

A male cone of Cycas circinalis

Selected species

Cycas aculeata Cycas angulata Cycas annaikalensis Cycas apoa Cycas arenicola Cycas armstrongii Cycas arnhemica Cycas badensis Cycas balansae Cycas basaltica Cycas beddomei Cycas bifida Cycas bougainvilleana Cycas brachycantha Cycas brunnea Cycas cairnsiana Cycas calcicola Cycas campestris Cycas candida Cycas canalis Cycas chamaoensis Cycas changjiangensis Cycas chevalieri Cycas circinalis Cycas clivicola Cycas collina Cycas condaoensis Cycas conferta Cycas couttsiana Cycas curranii Cycas debaoensis Cycas desolata Cycas diannanensis

Cycas dolichophylla Cycas edentata Cycas elephantipes Cycas elongata Cycas falcata Cycas fairylakea Cycas ferruginea Cycas fugax Cycas furfuracea Cycas guizhouensis Cycas hainanensis Cycas hoabinhensis Cycas hongheensis Cycas inermis Cycas javana Cycas lanepoolei Cycas lindstromii Cycas litoralis Cycas maconochiei Cycas macrocarpa Cycas media Cycas megacarpa Cycas micholitzii Cycas micronesica Cycas multipinnata Cycas nathorstii Cycas nongnoochiae Cycas ophiolitica Cycas orientis Cycas pachypoda Cycas panzhihuaensis Cycas papuana

Cycas pectinata Cycas petraea Cycas platyphylla Cycas pranburiensis Cycas pruinosa Cycas revoluta Cycas riuminiana Cycas rumphii Miq. Cycas schumanniana Cycas scratchleyana Cycas seemannii A.Braun Cycas segmentifida Cycas semota Cycas sexseminifera Cycas siamensis Cycas silvestris Cycas simplicipinna Cycas spherica Cycas szechuanensis Cycas taitungensis Cycas taiwaniana Cycas tanqingii Cycas tansachana Cycas thouarsii Cycas tropophylla Cycas tuckeri Cycas wadei Cycas xipholepis Cycas yorkiana Cycas yunnanensis Cycas zambalensis Cycas zeylanica

References and external links

• Hill KD (1998-2004) The Cycad Pages, Royal Botanic Gardens Sydney. http://plantnet.rbgsyd.nsw.gov.au/PlantNet/cycad/index.html
• Singh, R., & Radha P. (2006). Cycas annaikalensis, A new species of Cycas from the Malabar Coast, Western Ghats, India. Brittonia 58 (2): 119-123.
• Virtual Cycad Encyclopedia edited by the Palm & Cycad Societies of Florida
• David J. de Laubenfels, Cycas Taxonomy
• Ken D. Hill, Dennis W. Stevenson & Roy Osborne (2004) "The World List of Cycads". The Botanical Review, 70(2)274–298 doi:10.1663/0006-8101(2004)070%5B0274:TWLOC%5D2.0.CO;2
• Lindstrom, Anders J. and Ken D. Hill (2007) "The genus Cycas (Cycadaceae) in India". Telopea 11(4) : 463-488.
• David J. de Laubenfels & F. Adema (1998) "A taxonomic revision of the genera Cycas and Epicycas Gen. Nov. (Cycadaceae)", Blumea 43()351-400.
• Terrence Walters & Roy Osborne (eds.) (2004), Cycad Classification: Concepts and Recommendations, CABI publishing, ISBN 0-85199-741-4