Oecophorinae

Oecophorinae
Adult of Philobota arabella or a related species, Aranda (Australia)
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Lepidoptera
Suborder:	Glossata
Infraorder:	Heteroneura
Division:	Ditrysia
Superfamily:	Gelechioidea
Family:	Oecophoridae
Subfamily:	Oecophorinae Bruand, 1851
Genera
Numerous, see text
Synonyms
Chimabachidae Heinemann, 1870 Chimabachinae Deuterogoniidae Deuterogoniinae Enicostominae Heslop, 1938 Peleopodidae Peleopodinae Philobotidae Philobotinae (but see text)

The Oecophorinae are the nominate subfamily of moths in the concealer moth family (Oecophoridae). They are part of the insufficiently studied superfamily Gelechioidea, and like their relatives, the circumscription of this taxon is disputed.^[1]

In the approach followed here, the Oecophoridae are expanded to include several lineages formerly placed in the Elachistidae or considered independent gelechioid families. As regards the Oecophorinae, the proposed concealer moth subfamilies Chimabachinae, Deuterogoniinae, Peleopodinae and Philobotinae are included here pending further study of the affiliations of their genera. They were also often treated as independent families (Chimabachidae, Deuterogoniidae, Peleopodidae and Philobotidae) by those who followed a "splitting" approach. In general, the delimitation of the Oecophorinae versus the Amphisbatinae, Depressariinae and Hypertrophinae has been the most contested issue, though the uncertain placement of the Xyloryctidae versus the concealer moths (into which they might belong as subfamily) has also been a considerable stumbling block.^[1]

Numerous attempts have been made to divide the Oecophorinae into tribes, such as Carcinini, Crossotocerini, Denisiini, Herrichini, Oecophorini, Peleopodini and Pleurotini. Also placed here under this scheme are the Cacochroini and Orophiini, which otherwise were included in the Depressariinae (but usually only when these were elevated to full family rank). Most of the proposed tribes were based on phenetic or qualitative analyses, if not merely on the whim of the entomologists that established them, and no robust evolutionary scenario has been established for the different lineages of Oecophorinae. The groups around the genus Peleopoda (the former Peleopodinae) and of course the type genus Oecophora are generally recognized to be well distinguished from each other, but no satisfying arrangement has been found for the bulk of the (presumed) oecophorine genera. Hence, no subdivision into tribes is attempted here.^[1]

Selected genera

Adult of an unidentified Enchocrates species, Aranda (Australia)

Adult Garrha pudica, Aranda (Australia)

Adult of a newly-discovered Microbela species, Aranda (Australia)

Adult male Phaeosaces coarctatella, Browns Bay (New Zealand)

Adult of a newly-discovered Philobota species related to P. mathematica, Aranda (Australia)

Phryganeutis cinerea, Aranda (Australia)

Adult Pleurota aristella

The following genera (with some notable species also listed) are usually held to belong to the Oecophorinae, at least in the delimitation used here. Still, placement of few genera is completely certain (see above), and many – in particular monotypic genera – may not be valid at all. New oecophorine genera are also being described frequently:^[1]

Acanthodela Common, 1994 Acanthodela protophaes Acantholena Common, 1997 Acedesta Turner, 1940 Acmotoma Common, 1994 Acolasta Meyrick, 1902 Acorotricha Meyrick, 1913 Actenotis Turner, 1935 Aeolothapsa Common, 1997 Aeolothapsa malacella Ageletha Common, 1994 Ageletha hemiteles – Webbing Moth Aglaodes Turner, 1898 Airogephyra Common, 2000 Alabonia Allognoma Common, 1997 Analcodes Turner, 1947 Ancistromorpha Common, 1994 Ancistroneura Turner, 1947 Anomozancla Turner, 1936 Anthocoma Turner, 1946 Antiopala Meyrick, 1889 Antipterna Meyrick, 1916 Aplota Arachnographa Meyrick, 1914 Archaereta Meyrick, 1914 Aristeis Meyrick, 1883 Artiastis Meyrick, 1889 Ascetoloba Common, 1997 Ashinaga Matsumura, 1929 (previously in monotypic family Ashinagaidae) Aspasiodes Turner, 1944 Ataleida Common, 2000 Atalopsis Common, 2000 Atelosticha Meyrick, 1883 Atheropla Meyrick, 1883 Atheropla decaspila Atholosticta Common, 1997 Baiocystis Common, 1994 Baioglossa Common, 2000 Barea Walker, 1864 Barea codrella – Barea Moth Barea consignatella Barea melanodelta Basiplecta Common, 1994 Bathrosterra Common, 1994 Batia Stephens, 1834 Batia lambdella Batia lunaris – Lesser Tawny Crescent Batia unitella Bisigna Toll, 1956[verification needed] Bisigna procerella Borkhausenia Boroscena Common, 1997 Brachynemata Meyrick, 1883 Brymblia Hodges, 1974 Buvatina Leraut, 1984 Cacochroa Heinemann, 1870 Cacochroa permixtella Callimima Turner, 1935 Carcina Hübner, 1825[verification needed] Carcina quercana Carolana Clarke, 1941 Casmara Walker, 1863 Catacometes Common, 1994 Catadoceta Common, 2000 Cephalispheira Bruand, 1850[verification needed] Chezala Walker, 1864 Chioneocephala Common, 2000 Chrysonoma Meyrick, 1914 Cirrograpta Common, 2000 Cirromitra Common, 2000 Clonitica Meyrick, 1914 Cnecophora Common, 1996 Coelognatha Common, 2000 Coeranica Meyrick, 1883 Coesyra Meyrick, 1883 Compsotropha Meyrick, 1883 Compsotropha selenias Conobrosis Common, 1997 Conobrosis acervata Copidostola Lower, 1897 Corethropalpa Turner, 1896 Corocosma Meyrick, 1927[verification needed] Corocosma memorabilis Corynotricha Common, 2000 Coryphoscola Common, 1994 Cosmaresta Common, 1994 Cosmaresta charaxias Crassa Crepidosceles Meyrick, 1883 Crepidosceles exanthema Crossophora Crossotocera Zerny, 1930 Cryptotypa Common, 1997 Cyphacma Meyrick, 1915 Dasystoma Curtis 1833 Dasystoma salicella – Blueberry Leafroller Decantha Busck, 1908 Delexocha Common, 1997[verification needed] Delexocha ochrocausta Delophanes Turner, 1947 Denisia Deuterogonia Rebel 1901 Deigmoesta Common, 1994 Diapatela Common, 1997 Diaphanta Common, 1996 Diaphorodes Turner, 1946 Diasceta Common, 2000 Diocrogephyra Common, 2000 Dissoloba Common, 1997 Diplogephyra Common, 1997 Diplogrypa Common, 1997 Disselia Meyrick, 1883 Diurnea Haworth 1811 Diurnea fagella Diurnea lipsiella Dolopsis Common, 2000 Drepanocera Common, 1997 Durrantia Busck, 1908 Dysthreneta Turner, 1947 Echinobasis Common, 1994 Echinocosma Common, 1997 Echinognatha Common, 2000 Eido Chambers, 1875 Elaeonoma Meyrick, 1914 Elaphromorpha Turner, 1936 Enchocrates Meyrick, 1883 Enchronista Meyrick, 1914 Endeolena Common, 1994 Endrosis – White-shouldered House Moth Enlopholepis Common, 2000 Enoplidia Common, 1994 Eochrois Meyrick, 1886 Epicallima Dyar, 1903 Epicallima formosella

Epicurica Meyrick, 1914 Epithymema Turner, 1914 Eratophyes Diakonoff, 1975 Ereiconastes Common, 2000 Eremnotypa Common, 1994 Eremnozona Common, 2000 Ericibdela Common, 1994[verification needed] Ericibdela delotis Ericrypsina Common, 1997 Eridolera Common, 2000 Erythrisa Common, 1997 Erythrobapta Common, 2000 Esperia Euchaetis Meyrick, 1883 Euchaetis rhizobola Eulachna Meyrick, 1883 Eulechria Meyrick, 1883 Eulechria encratodes Eulechria subpunctella Euphiltra Meyrick, 1883 Eusemocosma Common, 1997[verification needed] Eusemocosma pruinosa Euthictis Meyrick, 1914 Exarsia Meyrick, 1914 Fabiola Busck, 1908 Garrha Walker, 1866 Garrha limbata Garrha ocellifera Goidanichiana Agenjo, 1976 Guestia Meyrick, 1889 Gymnocoila Common, 1997 Habrochlanis Common, 2000 Habroscopa Meyrick, 1914 Hadrocheta Common, 1994 Hadrognatha Common, 1997 Haplodyta Meyrick, 1883 Hapaloteucha Meyrick, 1914 Harpella Heliocausta Meyrick, 1883 Heloscopa (formerly in Tineidae) Hemibela Turner, 1894 Hemibela callista Herrichia Staudinger, [1871] Hesperenoeca Common, 2000 Hesperoptila Meyrick, 1902 Heterochyta Meyrick, 1906 Heteroptolis Meyrick, 1914 Heteroteucha Common, 1994 Heteroteucha parvula Heteroteucha translatella Heterozyga Meyrick, 1883[verification needed] Heterozyga coppatias Hofmannophila – Brown House Moth Holoscolia Zeller, 1839 Hoplomorpha Turner, 1916 Hoplomorpha camelaea Hoplostega Meyrick, 1914 Horridopalpus Hannemann 1953 (formerly in Depressaria) Hybocrossa Turner, 1917 Idioglossa Walsingham, 1881 Idioxantha Common, 1994 Idiozancla Turner, 1936 Inga Busck, 1908 Inga sparsiciliella – Black-marked Inga Moth Ioptera Meyrick, 1883 Ironopolia Common, 1994 Ironopolia sobriella Ischnomorpha Turner, 1937 Isomoralla Common, 1997 Isomoralla eriscota Izatha Walker, 1864 Joonggoora T.P.Lucas, 1901 Kasyniana Vives, 1986 Lamproxantha Common, 1994 Lasiocosma Common, 2000 Laxonoma Meyrick, 1914 Leimmatonca Common, 2000 Leipochlida Common, 2000 Leistomorpha Meyrick, 1883 Lepidotarsa Meyrick, 1883 Leprocosma Common, 2000 Leptocopa Meyrick, 1918 Leptocroca Meyrick, 1883 Leucorhabda Common, 1994 Limothnes Turner, 1935 Linosticha Meyrick, 1883 Liocnema Turner, 1941 Liozancla Turner, 1919 Lonchoptena Common, 2000 Locheutis Meyrick, 1883 Lophopepla Turner, 1896 Machaeritis Meyrick, 1883 Machetis Meyrick, 1883 Macronemata Meyrick, 1883 Macrophara Turner, 1946 Martyringa Busck, 1902 Mathildana Clarke, 1941 Meioglossa Common, 2000 Melanoima Common, 2000 Mermeristis Meyrick, 1915 Merocroca Common, 1997[verification needed] Merocroca automima Metalampra Metaphrastis Meyrick, 1907 Micramicta Common, 2000 Microbela Meyrick, 1883 Microlocha Meyrick, 1914 Micropeteina Common, 2000 Mimobrachyoma Lower, 1902 Mionolena Common, 1994 Myrascia Common, 1977 Nemepeira Common, 2000 Neosigala Turner, 1917 Nephogenes Meyrick, 1883 Ochlogenes Meyrick, 1883 Ochropolia Common, 2000 Ochyrolopha Common, 2000 Ocystola Meyrick, 1885 Ocystola paulinella Oecophora Oenochroa Meyrick, 1883 Olbonoma Meyrick, 1914 Olbonoma triptycha Olenacantha Common, 1997 Oligoloba Common, 1994 Oncolapara Common, 2000 Oncomerista Common, 2000 Opsitycha Meyrick, 1914 Orescoa Turner, 1927 Oresitropha Turner, 1927 Orophia Hübner 1825 Orthiastis Meyrick, 1914

Oxythecta Meyrick, 1883 Oxythecta acceptella Ozotrypetes Common, 2000 Pachybela Turner, 1917 Pachyceraia Common, 2000 Palimmeces Turner, 1916 Paneutricha Common, 1994 Pantogymna Common, 1997 Pararsia Turner, 1939 Parergophela Common, 1997 Parocystola Turner, 1896 Parocystola acroxantha – Ruddy Streak Pauronota Lower, 1901 Pedois Lower, 1894 Pedois humerana Pedois lewinella Pelinoema Common, 1997 Pellopsis Common, 1997 Periallactis Meyrick, 1902 Periorycta Meyrick, 1922 Petalanthes Meyrick, 1883 Phaeosaces Meyrick, 1885 Phauloglossa Common, 1997 Phanerozancla Turner, 1935 Phauloplana Common, 2000 Philobota Meyrick, 1883 Philobota lysizona Philobota partitella Philobota stella Phloeocetis Turner, 1936 Phloioletes Common, 2000 Pholeutis Meyrick, 1906 Phriconyma Meyrick, 1883 Phryganeutis Meyrick, 1884 Phryganeutis cinerea Phyllophanes Turner, 1896 Phytotrypa Common, 1994 Phyzanica Turner, 1917 Piloprepes Meyrick, 1883 Piloprepes anassa Piloprepes antidoxa Placocosma Meyrick, 1883 Placocosma resumptella Platoloncha Common, 1997 Platyphylla Turner, 1946 Plectobela Common, 1994 Plectobela zanclotoma Pleurota Hübner, 1825 Pleurota bicostella Poliorhabda Common, 1994 Polix Hodges, 1974 Polyeucta Turner, 1917 Prepalla Common, 1997 Prepocosma Common, 1997 Prionocris Common, 1994 Prodelaca Common, 1994 Prodelodes Common, 1997 Proteromicta Meyrick, 1889 Protomacha Meyrick, 1883 Psaltriodes Meyrick, 1902 Psarophorca Common, 2000 Psaroxantha Common, 1994 Psaroxantha calligenes Pseudocryptolechia Lvovky, 2001 Pseudoecophora Staudinger, 1899 Pseudotheta Clarke, 1947 Ptochosaris Meyrick, 1906 Ptyoptila Turner, 1946[verification needed] Ptyoptila matutinella Pycnocera Turner, 1896 Pycnozancla Turner, 1917 Pyrgoptila Meyrick, 1889 Rhadinoloba Common, 1994 Rhoecoceros Turner, 1940 Saphezona Common, 1994 Saropla Meyrick, 1883 Satrapia Meyrick, 1883 Scatochresis Common, 1997 Schiffermuelleria Sclerocheta Common, 1994 Sclerocris Common, 1997 Scoliocheta Common, 2000 Scotodryas Turner, 1932 Sphyrelata Meyrick, 1883 Stenoptena Common, 2000 Stereocheta Common, 1997 Stereoloba Common, 1994 Sthenozancla Turner, 1941 Stictochila Common, 1994 Stictopolia Common, 1997 Sympoecila Common, 2000 Syncometes Common, 2000 Syringoseca Common, 1994 Syringoseca rhodoxantha Syscalma Meyrick, 1920 Tachystola Meyrick, 1914[verification needed] Tachystola acroxantha Tanycaula Common, 1997 Tanyzancla Meyrick, 1918 Teerahna T.P.Lucas, 1901 Telanepsia Turner, 1933 Telocharacta Common, 1997 Temnogyropa Common, 1997 Temnogyropa stenomorpha Thalerotricha Meyrick, 1883 Thapsinotypa Common, 2000 Thema Walker, 1864 Thema macroscia Thema protogramma Thema psammoxantha Tingena Walker, 1864 Tingena armigerella Tingena marcida Tisobarica Walker, 1864 Tisobarica thyteria Tortricopsis Newman, 1856 Tortricopsis euryphanella Tortricopsis pyroptis Tortricopsis semijunctella Tortricopsis uncinella Trachyxysta Meyrick, 1916 Trachyzancla Turner, 1917 Triacra Common, 2000 Trichomoeris Meyrick, 1913 Trinaconeura Turner, 1933 Wingia Walsingham & Durrant, 1900 Wingia aurata – Golden Leaf Moth Woorda T.P.Lucas, 1901 Wullaburra T.P.Lucas, 1901 Zacorus Butler, 1882 Zelotechna Meyrick, 1914 Zonopetala Meyrick, 1883 Zonopetala clerota Zonopetala decisana

Several genera of Oecophoridae are of very unclear or highly disputed relationships. Among these, some have been theorized to belong in the Oecophoridae:

• Callimodes Leraut 1989
• Minetia Leraut, 1991
• Schiffermuellerina Leraut, 1989^{[verification needed]}
  • Schiffermuellerina grandis
• Zizyphia (formerly in Gelechiidae)

Also, it may be that some additional genera usually placed outside the Gelechioidea are actually Oecophorniae, too, e.g.:

• Leurophanes Turner, 1939 (usually in Plutellidae)
• Notodryas Meyrick, 1897 (usually in Epermeniidae)
• Trisyntopa Lower, 1918 (usually in Tineidae)

Footnotes

1. ^ ^{a b c d} ABRS (2008), FE (2009), Wikispecies (2010-APR-25), and see references in Savela (2009)

References

Data related to Oecophorinae at Wikispecies See also Gelechioidea Talk page for comparison of some approaches to gelechioid systematics and taxonomy.

• Australian Biological Resources Study (ABRS) (2008): Australian Faunal Directory – Oecophorinae. Version of 2008-OCT-09. Retrieved 2010-APR-24.
• Fauna Europaea (FE) (2009): Gelechioidea. Version 2.1, 2009-DEC-22. Retrieved 2010-APR-22.
• Savela, Markku (2009): Markku Savela's Lepidoptera and some other life forms – Oecophorinae. Version of 2009-AUG-20. Retrieved 2010-APR-22.