Microberotha

Microberotha macculloughi Temporal range: Ypresian PreЄ Є O S D C P T J K Pg N
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Neuroptera
Suborder:	Hemerobiiformia
Superfamily:	Mantispoidea
Family:	Berothidae
Subfamily:	incertae sedis
Genus:	† Microberotha
Species:	M. macculloughi Archibald & Makarkin, 2004
Binomial name
Microberotha macculloughi Archibald & Makarkin, 2004

Microberotha is an extinct monotypic genus of "beaded lacewing", family Berothidae; order Neuroptera with the single species Microberotha macculloughi.^[1](p61) The genus is known from a single male specimen, the holotype, currently deposited in the collections of the Kelowna Centennial Museum in Kelowna, British Columbia, as number "NH.998.015.002", and which was first studied by Dr Bruce Archibald and Dr Vladimir Makarkin. Drs Archibald and Makarkin published their type description in the journal The Canadian Entomologist volume 136 published in 2004.^[1](p61) The genus name is a combination of the Greek mikros meaning "small" and Berotha, a genus in the family Berothidae. The species name is in honor of Terry McCullough, a supporter of paleontological research at the type locality.^[1](p66) "Mac" is used in the species epithet to conform to the International Code of Zoological Nomenclature recommendations regarding names from personal names.^[1](p66) The type specimen is well preserved in early Early Eocene^[1](p64) Hat Creek amber from deposits between Cache Creek and Lillooet, British Columbia that are part of the greater Okanagan Highlands paleoenvironment.^[1](p63)

Microberotha can be distinguished from other berothids by the combination of an unbranched "Rs" vein and lack of an outer gradate series of crossveins in the forewings.^[1](p65) With a forewing length of only 3.36 millimetres (0.132 in) M. macculloughi is one of the smallest known members of Berothidae to have been described.^[1](p70) The smallest living species Manselliberotha neuropterologorum has a minimum forewing length of 3.6 millimetres (0.14 in), while some species from the New Jersey ambers are smaller at a minimum of 2.6 millimetres (0.10 in).^[1](p70-71) The terminal segments of the abdomen on the holotype, necessary in determining subfamily placement, are not visible given the angles of view available through the amber.^[1](p71) This difficulty prevents the genus from being placed at the subfamily level and thus is incertae sedis in the family.^[1](p71-72) The holotype is preserved in a specimen of amber which shows distinct flow patterning indicating it started as a "stalactite" or similar formation on the tree.^[1](p65) The positioning of the insect relative to the flow pattern indicates it was most likely trapped with its back stuck to the inner flow with its left side facing down.^[1](p65) A later thicker flow covered the insect twisting the left hind wing and disarticulating both the hind legs.^[1](p65)

References

1. ^ ^{a b c d e f g h i j k l m n} Archibald, S.B.; Makarkin, V.N. (2004). "New genus of minute Berothidae (Neuroptera) from Early Eocene amber of British Columbia". The Canadian Entomologist 136 (1): 61–76. doi:10.4039/n03-043. http://ibss.febras.ru/files/00002082.pdf.