Ophthalmosauridae

Ophthalmosauridae
Ophthalmosauridae
Temporal range: Middle Jurassic-Late Cretaceous, 171.6–93.5 Ma
Ophthalmosaurus icenicus skeleton
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Ichthyosauria
Node: Ophthalmosauria
Family: Ophthalmosauridae
Baur, 1887
Genera

Ophthalmosauridae is an extinct family of thunnosaur ichthyosaurs from the Middle Jurassic to the early Late Cretaceous (Bajocian - Cenomanian) of Asia, Australia, Europe, North America and South America.[1] Currently, the oldest known ophthalmosaurid is Mollesaurus from the early Bajocian of Argentina.[2] Named by George H. Baur, in 1887, it contains the basal taxa like Ophthalmosaurus.[1] Appleby (1956) named the taxon Ophthalmosauria which was followed by some authors,[3][4] but if these two names are synonyms, Ophthalmosauridae has the priority over Ophthalmosauria.[1]

Phylogeny

Ophthalmosaurus

Ophthalmosauridae is a node-based taxon defined by Ryosuke Motani (1999) as "the last common ancestor of Brachypterygius extremus and Ophthalmosaurus icenicus and all of its descendants". The definition he proposed for Ophthalmosauria was exactly the same.[3] In this case both defintions are synonyms. The cladogram below follows the topology from a 2010 analysis by Patrick S. Druckenmiller and Erin E. Maxwell.[4]

Thunnosauria 

Ichthyosaurus




Stenopterygius




"Ophthalmosaurus" natans


 Ophthalmosauridae 


Aegirosaurus



Ophthalmosaurus (type species)





Mollesaurus




Athabascasaurus




Brachypterygius




Arthropterygius



Caypullisaurus



"Platypterygius" hercynicus



"Platypterygius" australis



Platypterygius (type species)




Maiaspondylus



"Platypterygius" americanus











Valentin Fischer, Edwige Masure, Maxim S. Arkhangelsky and Pascal Godefroit (2011) described a new genus of Ophthalmosauridae. They redefined it as "the last common ancestor of Arthropterygius chrisorum and Ophthalmosaurus icenicus and all of its descendants". In this case Ophthalmosauria isn't a junior synonym of Ophthalmosauridae. The cladogram below follows Fischer et al. 2011.[1]

Thunnosauria 

Ichthyosaurus




Stenopterygius




Chacaicosaurus


 Ophthalmosauridae 

Arthropterygius


*

Brachypterygius



Ophthalmosaurus (type species)





Maiaspondylus




Mollesaurus




Aegirosaurus



Sveltonectes







"Platypterygius" hercynicus




Caypullisaurus




Athabascasaurus



"Platypterygius" australis











*Note: Placement of Ophthalmosauria by definition.

References