Orthalicidae

Orthalicidae
A large orthalicid from eastern Peru (probably of the genus Sultana), recovering from substantial shell damage
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
(unranked):	clade Heterobranchia clade Euthyneura clade Panpulmonata clade Eupulmonata clade Stylommatophora informal group Sigmurethra
Superfamily:	Orthalicoidea
Family:	Orthalicidae Albers, 1860[1]
Subfamilies
Amphibuliminae P.Fischer, 1873 Bulimulinae Tryon, 1867 Orthalicinae Albers, 1850
Diversity
~1450 species (without Amphibulimidae)[2]
Synonyms
Bulimulidae Crosse & P. Fischer, 1873 (but see text)

Orthalicidae (orthalicid land snails) are a family of tropical air-breathing land snails, terrestrial pulmonate gastropod mollusks are classified in the informal group Sigmurethra of the clade Stylommatophora. They are medium-sized to large snails, from about 3 cm (about 1.2 inches) to 9 cm (about 3.5 inches) in shell length.

Taxonomy

In former times, this family was often known as the Bulimulidae, but this term may also denote what today is the subfamily Bulimulinae. The subfamily Bulimulinae replaces the former family Bulimulidae Crosse & P. Fischer, 1873.

2005 taxonomy

The Orthalicidae belong to the Orthalicoidea, a superfamily in the clade Sigmurethra. Like other stylommatophorans, the Sigmurethra belong to the suborder Eupulmonata. Among the three subfamilies of Orthalicidae, the Bulimulinae are the most diverse, consisting of three tribes:^[3]

• subfamily Amphibuliminae P. Fischer, 1873 - synonym Peltellinae Gray, 1855 (check decision of International Commission on Zoological Nomenclature for Peltellinae) - 17 species^[2]
• subfamily Bulimulinae Tryon, 1867 - 1243 species^[2]
  • tribe Bulimulini Tryon, 1867 - synonyms: Bulimidae Guilding, 1828 (inv.); Berendtiinae P. Fischer & Crosse, 1872; Bothiembryontidae Iredale, 1937
  • tribe Odontostomini Pilsbry & Vanatta, 1898 - 125 species^[2]
  • tribe Simpulopsini Schileyko, 1999 - synonym: Tomogeridae Jousseaume, 1877
• subfamily Orthalicinae Albers, 1850 - synonym: Liguidae Pilsbry, 1891 - 83 species^[2]

Genera

Genera within the family Orthalicidae include:

subfamily Amphibuliminae

• Amphibulima Lamarck, 1805 - type genus of the subfamily Amphibuliminae^[3]
• Peltella Gray, 1855

subfamily Bulimulinae

Main article: Bulimulinae

• Bulimulus Leach, 1814 - type genus of the subfamily Bulimulinae^[3]

subfamily Orthalicinae

• Corona Albers, 1850^[4][5]
• Hemibulimus Martens, 1885^[4]
• Porphyrobaphe Shuttleworth, 1856^[4]
• Liguus Montfort, 1810
• Orthalicus Beck, 1837 - type genus of the family Orthalicidae^[3]
• Sultana Shuttleworth, 1856^[4]

References

1. ^ Albers J. C. 1860. (Manuscript edited by Eduard von Martens) Die Heliceeen nach natürlicher Vervandtschaft systematisch geordnet. Edition 2, Engelmann, Leipzig, xviii + 359 pp., Orthalicidae is at page 209.
2. ^ ^{a b c d e} Breure A., Groenenberg D. & Schilthuizen M. (2010). "Gondwana revisited: New insights in the phylogenetic relationships within the Orthalicoidea (Mollusca, Stylommatophora)". poster. PDF.
3. ^ ^{a b c d} Bouchet P., Rocroi J.-P., Frýda J., Hausdorf B., Ponder W., Valdés Á. & Warén A. (2005). "Classification and nomenclator of gastropod families". Malacologia: International Journal of Malacology (Hackenheim, Germany: ConchBooks) 47 (1-2): 1–397. ISBN 3925919724. ISSN 0076-2997. http://www.archive.org/details/malacologia47122005inst. 
4. ^ ^{a b c d} Mónica Lucía Vera Ardila (2008). "Lista de los géneros de moluscos terrestres de Colombia (Mollusca: Gastropoda: Prosobranchia: Mesogastropoda y Pulmonata: Stylommatophora)". Biota Colombiana 9 (1): 39–62. http://www.siac.net.co/biota/bitstream/123456789/245/1/malacofauna.pdf. 
5. ^ Breure A. S. H. & Mogollón Avila V. (2010). "Well-known and little-known: miscellaneous notes on Peruvian Orthalicidae (Gastropoda, Stylommatophora)". Zoologische Mededelingen 84. HTM.

Further reading

• (German) Neubert E. & Janssen R. (2004). "Die Typen und Typoide des Natur-Museums Senckenberg, 84: Mollusca: Gastropoda: Pulmonata: Orthalicoidea: Bulimulidae (2), Orthalicidae, Placostylidae". Archiv für Molluskenkunde 133: 193-297. abstract.
• Ramirez J. & Ramírez R. (2010). "Análisis de la estructura secundaria del LSU rRNA mitocondrial de caracoles terrestres peruanos (Orthalicidae: Gastropoda) / Analysis of the secondary structure of mitochondrial LSU rRNA of Peruvian land snails (Orthalicidae: Gastropoda)." Revista Peruana de Biología 17(1): 53-57. PDF.
• Ramirez J., Ramírez R., Romero P., Chumbe A. & Ramírez P. (2009). "Posición evolutiva de caracoles terrestres peruanos (Orthalicidae) entre los Stylommatophora (Mollusca: Gastropoda) / Evolutionary position of Peruvian land snails (Orthalicidae) among Stylommatophora (Mollusca: Gastropoda)". Revista Peruana de Biología 16(1): 51-56. PDF.

External links

• Orthalicidae at National Center for Biotechnology Information (NCBI) website.