Elachista

For the genus of Chordariaceae brown algae, see Elachista (alga). As invalidly described by V. Kollar in 1832, Elachista refers to the Lyonetiidae moth genus Lyonetia.

Elachista
Adult Elachista argentella at IJmuiden (The Netherlands)
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Lepidoptera
Division:	Ditrysia
Family:	Elachistidae
Genus:	Elachista Treitschke in Ochsenheimer, 1833
Type species
Elachista bifasciella Treitschke in Ochsenheimer, 1833
Species
Numerous, see text
Synonyms
Numerous, see text

Elachista is a genus of gelechioid moths. It is the type genus of the grass-miner moth family (Elachistidae). This family is sometimes (in particular in older sources) circumscribed very loosely, including for example the Agonoxenidae and Ethmiidae which seem to be quite distinct among the Gelechioidea, as well as other lineages which are widely held to be closer to Oecophora than to Elachista and are thus placed in the concealer moth family Oecophoridae here.^[1]

These grass-miners are very small moths with the "feathery" hindwings characteristic of their family. They are essentially found worldwide, except in very cold places and on some oceanic islands; as usual for Gelechioidea, they are most common in the Palearctic however. They usually have at least one, sometimes as many as three light bands running from leading to trailing edge of their forewing uppersides. Some species, however, have upper forewings that are mostly white.^[2]

Selected species

Not all species of this large genus have been discovered yet, let alone validly described and named. Several small genera, e.g. Biselachista and Cosmiotes, are here included in Elachista as junior synonyms (see below). They seem to be at least as closely related to E. bifasciella (the type species of the present genus) than other species commonly placed here, if not actually closer. In addition, Elachista contains several cryptic species complexes – such as the one around E. dispunctella and E. triseriatella – whose systematics and taxonomy are still not fully resolved.^[3]

A number of Elachista species have been assigned to one of several groups, which may or may not be monophyletic, and whose names in the literature do not consistently follow the usual taxonomic practice (i.e. using as namesake the group-member which was described first). Some of these groups are placed in either of the two large subgenera Elachista (Aphelosetia) and Elachista (Elachista). Other proposed subgenera – the small Elachista (Dibrachia) and Elachista (Hemiprosopa) – are here included in the species of unclear relationships; like some small species-groups, they stand a chance of rendering the two larger subgenera paraphyletic if accepted as distinct, but seem too unlike them to warrant placement in either.^[4]

Subgenus Elachista (Aphelosetia)

argentella group (sometimes in bedellella group)

• Elachista argentella (type of Aphelosetia)
• Elachista habeleri Traugott-Olsen, 1990
• Elachista ripula Kaila, 1998
• Elachista triatomea
• Elachista virgatula Kaila, 1997 (type of Aphigalia)

Adult male Elachista pullicomella specimen from Tvärminne (Hanko Peninsula, Finland)

bedellella/unifasciella group (including collitella group)

Elachista adscitella Stainton, 1851 Elachista antonia Kaila, 2007 Elachista bedellella (Sircom, 1848) Elachista bisulcella (Duponchel, [1843]) Elachista bruuni Traugott-Olsen, 1990 Elachista camilla Kaila, 2007 Elachista cahorsensis Traugott-Olsen, 1992 Elachista chrysodesmella Zeller, 1850 Elachista cingillella (Herrich-Schäffer, 1855) Elachista collitella (Duponchel, [1843]) Elachista crocogastra Meyrick, 1908 Elachista dispilella Zeller, 1839 Elachista dispunctella (Duponchel, [1843]) Elachista distigmatella Frey, 1859 Elachista dorinda Kaila, 2007 Elachista fasciola Parenti, 1983 Elachista festucicolella Zeller, 1853 Elachista gangabella Zeller, 1850

Elachista gregori Traugott-Olsen, 1988 Elachista heinemanni[verification needed] Frey, 1866 (sometimes in E. subalbidella; tentatively placed here) Elachista istanella Nielsen[verification needed] & Traugott-Olsen, 1987 Elachista laetella Rebel, 1930 (sometimes in E. subalbidella; tentatively placed here) Elachista levasi Sruoga, 1998 Elachista littoricola Le Marchand, 1938 Elachista lugdunensis Frey, 1859 Elachista metella Kaila, 2002 Elachista nedaella Traugott-Olsen, 1985 Elachista nitidulella (Herrich-Schäffer, 1855) Elachista nolckeni Sulcs, 1992 (tentatively placed here) Elachista obliquella Stainton, 1854 Elachista pollinariella Zeller, 1839

Elachista pollutissima Staudinger, 1880 Elachista pullicomella Elachista purella Sruoga, 2000 Elachista revinctella Zeller, 1850 Elachista rudectella Stainton, 1851 Elachista slivenica Kaila, 2007 Elachista squamosella (Duponchel, [1843]) Elachista subalbidella Elachista subocellea Elachista sutteri Kaila, 2002 Elachista tinctella Sinev & Sruoga, 1995 Elachista titanella Kaila & Jalava, 1994 Elachista triseriatella Stainton, 1854 Elachista unifasciella (Haworth, 1828) Elachista versicolora Kaila, 2007 Elachista acutella Kaila, 2003 Elachista agelensis Traugott-Olsen, 1996 Elachista amparoae Traugott-Olsen, 1992 Elachista amseli Rebel, 1833 Elachista andorraensis Traugott-Olsen, 1988 Elachista anitella Traugott-Olsen, 1985 Elachista arduella Kaila, 2003 Elachista atrisquamosa Staudinger, 1880 Elachista baldizzonei Traugott-Olsen, 1996 Elachista baldizzonella Traugott-Olsen, 1985 Elachista bazaella Traugott-Olsen, 1992 Elachista bazaensis Traugott-Olsen, 1990 Elachista bengtssoni Traugott-Olsen, 1992 Elachista berndtiella Traugott-Olsen, 1985 Elachista bigorrensis Traugott-Olsen, 1990 Elachista casascoensis Traugott-Olsen, 1992 Elachista catalana Parenti, 1978 Elachista catalunella Traugott-Olsen, 1992 Elachista chamaea Kaila, 2003 Elachista chionella Mann, 1861 Elachista clintoni Traugott-Olsen, 1992 Elachista constitella Frey, 1859 Elachista contisella Chrétien, 1922 Elachista cuencaensis Traugott-Olsen, 1992 Elachista curonensis Traugott-Olsen, 1990 Elachista dalmatiensis Traugott-Olsen, 1992 Elachista deceptricula Staudinger, 1880 Elachista disemiella Zeller, 1847 Elachista drenovoi Parenti, 1981 Elachista dumosa Parenti, 1981 Elachista elsaella Traugott-Olsen, 1988

Elachista exigua Parenti, 1978 Elachista flavescens Parenti, 1981 Elachista fuscibasella Chrétien, 1915 Elachista galacticella Eversmann, 1844 Elachista gerdmaritella Traugott-Olsen, 1992 Elachista gibbera Kaila, 2003 Elachista gielisi Traugott-Olsen, 1992 Elachista glaseri Traugott-Olsen, 1992 Elachista gormella Nielsen[verification needed] & Traugott-Olsen, 1987 Elachista graeca Parenti, 2002 Elachista grandella Traugott-Olsen, 1992 Elachista hallini Traugott-Olsen, 1992 Elachista hedemanni Rebel, 1899 Elachista heringi Rebel, 1899 Elachista hispanica Traugott-Olsen, 1992 Elachista imbi Traugott-Olsen, 1992 Elachista intrigella Traugott-Olsen, 1992 Elachista jaeckhi Traugott-Olsen, 1990 Elachista karsholti Traugott-Olsen, 1992 Elachista klimeschiella Parenti, 2002 Elachista latipenella Sinev & Budashkin, 1991 Elachista lerauti Traugott-Olsen, 1992 Elachista liskai Kaila, 2011 Elachista louisae Traugott-Olsen, 1992 Elachista luqueti Traugott-Olsen, 1992 Elachista maculata Parenti, 1978 Elachista madridensis Traugott-Olsen, 1992 Elachista mannella Traugott-Olsen, 1992 Elachista manni Traugott-Olsen, 1990 Elachista minusculella Traugott-Olsen, 1992 Elachista modesta Parenti, 1978 Elachista multipunctella Traugott-Olsen, 1992

Elachista neapolisella Traugott-Olsen, 1985 Elachista nevadensis Parenti, 1978 Elachista nielspederi Traugott-Olsen, 1992 Elachista nuraghella Amsel, 1951 Elachista occidentella Traugott-Olsen, 1992 Elachista occulta Parenti, 1978 Elachista ohridella Parenti, 2001 Elachista olschwangi Kaila, 2003 Elachista ozeini Parenti, 2004 Elachista parvula Parenti, 1978 Elachista passerini Traugott-Olsen, 1996 Elachista pocopunctella Traugott-Olsen, 1992 Elachista pollutella Duponchel, 1843 Elachista povolnyi Traugott-Olsen, 1992 Elachista punctella Traugott-Olsen, 1992 Elachista rikkeae Traugott-Olsen, 1992 Elachista skulei Traugott-Olsen, 1992 Elachista spumella Caradja, 1920 Elachista steueri Traugott-Olsen, 1990 Elachista svenssoni Traugott-Olsen, 1988 Elachista szocsi Parenti, 1978 Elachista teruelensis Traugott-Olsen, 1990 Elachista totanaensis Traugott-Olsen, 1992 Elachista toveella Traugott-Olsen, 1985 Elachista tribertiella Traugott-Olsen, 1985 Elachista vanderwolfi Traugott-Olsen, 1992 Elachista varensis Traugott-Olsen, 1992 Elachista vegliae Parenti, 1978 Elachista veletaella Traugott-Olsen, 1992 Elachista vivesi Traugott-Olsen, 1992 Elachista vulcana Kaila, 2011

Subgenus Elachista (Atachia)

These are sorted in the presumed phylogenetic sequence.^[6]

gerasmia group, gerasmia section

Incertae sedis Elachista magidina Elachista floccella Elachista velutina Elachista carcharota Elachista crenatella Elachista bidens Elachista alampeta Elachista cursa Elachista eriodes Elachista illota Elachista ceratiola Elachista bilobella Elachista filiphila Elachista merista Elachista lopadina Elachista patania Elachista indigens Elachista averta Elachista anolba Elachista flavicilia Elachista tetraquetri Elachista effusi Elachista deusta Elachista rubiginosae Elachista ophelma Elachista catagma Elachista ruscella Elachista sarota Elachista syntomella Elachista scopulina Elachista nodosae Elachista discina

Elachista cynopa complex Elachista ligula Elachista toryna Elachista ascidiella Elachista repanda Elachista prolatella Elachista protensa Elachista campsella Elachista spinodora Elachista glossina Elachista opima Elachista pharetra Elachista rudicula Elachista corbicula Elachista mystropa Elachista trulla Elachista euthema Elachista cynopa Elachista epartica Elachista zeta Elachista chilotera Elachista mundula Elachista listrionea Elachista crumilla Elachista aposematica Elachista chloropepla

Elachista melanthes complex Elachista melanthes Elachista platysma Elachista stictifica Elachista lachnella Elachista gerasmia Elachista physalodes Elachista peridiola Elachista phascola Elachista paragauda complex Elachista paragauda Elachista toralis Elachista paryphoea Elachista spathacea Elachista cylistica Elachista fabarella Elachista etorella Elachista gladiograpta Elachista gladiatrix Elachista spongicola

gerasmia group, catarata section

Elachista menura Elachista flammula Elachista crocospila Elachista glomerella Elachista sphaerella Elachista sapphirella

Elachista patersoniae Kaila, 2011 Elachista catarata Meyrick, 1897 Elachista mutarata Kaila, 2011 Elachista laterina Kaila, 2011 Elachista ictera Kaila, 2011 Elachista asperae Kaila, 2011 Elachista ophthalma Kaila, 2011 Elachista coalita Kaila, 2011 Elachista corneola Kaila, 2011

erebophthalma group

• Elachista erebophthalma
• Elachista evexa
• Elachista rhomboidea
• Elachista leucastra

Subgenus Elachista (Elachista)

Adult female Elachista apicipunctella specimen from Sonkajärvi (Finland)

Adult Elachista maculicerusella specimen from Hellerup (Denmark)

Adult male Elachista gleichenella specimen from Kangasniemi (Finland)

Adult male Elachista humilis specimen from Sonkajärvi (Finland)

albifrontella/bifasciella group Elachista albifrontella (Hübner, [1817]) Elachista alpinella Elachista apicipunctella Elachista atricomella Elachista baikalica Kaila, 1992 Elachista bifasciella Treitschke, 1833 Elachista compsa Traugott-Olsen, 1974 Elachista deriventa Kaila & Mutanen, 2008 Elachista diederichsiella E.Hering, 1889 Elachista elegans Frey, 1859 Elachista excelsicola Elachista fuscofrontella Sruoga, 1990 Elachista habrella Elachista hiranoi Sugisima, 2005 Elachista jaskai Kaila, 1998 Elachista jupiter Sugisima, 2005 Elachista kilmunella Stainton, 1849 Elachista leifi Kaila & Kerppola, 1992 Elachista luticomella Elachista micalis Elachista nobilella Elachista phalaridis Parenti, 1983 Elachista platina Elachista poae Stainton, 1855 Elachista scitula Elachista tanaella Aarvik & Berggren, 2004[verification needed] Elachista anserinella Zeller, 1839 Elachista lastrella Chrètien, 1896 Elachista maculicerusella Elachista rufocinerea Elachista wieseriella Huemer[verification needed], 2000 (tentatively placed here) gleichenella group Elachista enitescens Braun, 1921 Elachista gerasimovi Sruoga, 2000 Elachista gleichenella Elachista ievae Sruoga, 2008 Elachista madarella (Clemens, 1860) Elachista nitensella Sinev & Sruoga, 1995 Elachista regificella Sircom, 1849 Elachista similis Sugisima, 2005 Elachista tengstromi (tentatively placed here) orba group Elachista commoncommelinae Elachista cyanea Elachista polliae Elachista nielsencommelinae Elachista orba Meyrick 1921 Elachista amamaii Parenti, 1983 Elachista aurita Elachista caliginosa Parenti, 1983 Elachista fasciocaliginosa Sugisima, 2005 Elachista kurokoi Parenti, 1983 Elachista miscanthi Parenti, 1983 Elachista praelineata Braun 1915 pulchella group Elachista canapennella Elachista eskoi Elachista herrichii Frey, 1859 Elachista humilis Elachista krogeri Svensson, 1976 Elachista nielswolffi Svensson, 1976 Elachista orstadii Palm, 1943 Elachista pomerana Elachista subnigrella Douglas, 1853 Elachista vonschantzi Svensson, 1976 Elachista zernyi solena group Elachista ignicolor Elachista solena

Adult female Elachista exactella specimen from Sonkajärvi (Finland)

Adult male Elachista exactella specimen

Adult Elachista synethes specimen from ventral

"Cosmiotes"/freyerella group

Incertae sedis Elachista albipalpella Chambers, 1880[verification needed] Elachista albrechti Kaila, 1998 Elachista amseli (Parenti, 1981) Elachista antipetra Meyrick, 1922 Elachista baltica E.Hering, 1891 (sometimes in E. freyerella; tentatively placed here) Elachista consortella Stainton, 1851 (tentatively placed here) Elachista cristatella Chambers, 1876 Elachista deficiens Meyrick, 1922 Elachista epicaeria (Diakonoff, 1955) Elachista exactella (tentatively placed here) Elachista herbigrada Braun, 1925 Elachista heteroplaca Meyrick, 1934 Elachista illectella (Clemens, 1860) (type of Cosmiotes) Elachista illicitella[verification needed] (Clemens, 1860) Elachista infuscata Frey, 1882 (sometimes in E. exactella; tentatively placed here) Elachista inscia (Meyrick, 1913) Elachista ksarella Chrétien, 1908 Elachista laquaeorum (Dugdale, 1971) Elachista lorigera (Meyrick, 1921) (type of Ptilodoxa) Elachista nipponicella Sugisima, 2006 Elachista nymphaea Meyrick, 1911 Elachista obtusella Sruoga, 2008 Elachista ochroleuca Meyrick 1923 Elachista pallens (Sruoga, 1990) Elachista planicara Kaila, 1998 Elachista praematurella Clemens, 1860[verification needed] Elachista quadrata Meyrick, 1932 Elachista scopulicola (Braun, 1948) Elachista spiculifera Meyrick, 1922 Elachista stabilella Stainton, 1858 (tentatively placed here) Elachista stichospora Meyrick, 1932 Elachista tuberella Sruoga, 2008

Elachista freyerella complex[7] Elachista propera Elachista dieropa Elachista elaphria Elachista cycotis Elachista vitellina Elachista citrina Elachista cerebrosella Elachista melina Elachista gemadella Elachista ravella Elachista cerina Elachista diligens Elachista aepsera Elachista zophosema Elachista litharga Elachista levipes Elachista cerrita Elachista velox Elachista impiger Elachista alacera Elachista essymena Elachista festina

Elachista synethes complex[7] Elachista fucosa Elachista seductilis Elachista synethes Elachista strenua Elachista sandaraca Elachista helvola Elachista delira Adult female Elachista eleochariella specimen from Sonkajärvi (Finland) tetragonella group Elachista abiskoella Bengtsson, 1977 Elachista agilis Braun, 1921 Elachista albidella Elachista biatomella (Stainton, 1848) Elachista bilbaensis (Rebel, 1893) Elachista boursini Amsel, 1951 Elachista christenseni Traugott-Olsen, 2000 Elachista cinereopunctella (Haworth, 1828) Elachista cucullata Braun, 1921 Elachista eleochariella Elachista ensifera Elachista falirakiensis Traugott-Olsen, 2000 Elachista fulgens Parenti, 1983 (tentatively placed here) Elachista glaserella Traugott-Olsen, 2000 Elachista imatrella von Schantz, 1971 Elachista jordanella Amsel, 1935 Elachista juliensis Frey, 1870 = E. freyi Staudinger, 1871 (type of Biselachista) Elachista kebneella (Traugott-Olsen & Nielsen[verification needed], 1977) Elachista kleini Amsel, 1935 Elachista leucosticta Braun, 1948 Elachista leucosyrma (Meyrick, 1932) (type of Platyphyllis) Elachista martinii Hofmann, [1898] Elachista nevadella Traugott-Olsen, 2000 Elachista occidentalis Frey, 1882 Elachista ornithopodella Frey, 1859 Elachista pigerella (Herrich-Schäffer, 1854) (type of Atachia) Elachista quadripunctella (Hübner, [1825]) Elachista ribentella Kaila & Varalda, 2004 Elachista saarelai Elachista salinaris Braun, 1925 Elachista scirpi Stainton, 1887 Elachista serricornis Elachista tetragonella (Herrich-Schäffer, 1855) Elachista trapeziella Stainton, 1849 Elachista utonella Frey, 1856 Elachista zabella Chrétien, 1908 Elachista zonulae (Sruoga, 1992) Elachista adelpha Kaila & Jalava, 1994 Elachista albicapilla Höfner, 1918 Elachista anserinelloides Nel, 2003 Elachista argentifasciella Höfner, 1898 Elachista arnoldi (Koster, 1993) Elachista brachypterella (Klimesch, 1990) Elachista canariella Nielsen[verification needed] & Traugott-Olsen, 1987 Elachista contaminatella Zeller, 1847 Elachista devexella Kaila, 2003 Elachista differens Parenti, 1978 Elachista dimicatella Rebel, 1903 Elachista encumeadae Kaila & Karsholt, 2002 Elachista geminatella (Herrich-Schäffer, 1855) Elachista griseella (Duponchel, 1843) Elachista gruenewaldi Parenti, 2002 Elachista ibericella Traugott-Olsen, 1995 Elachista igaloensis (Amsel, 1951) Elachista irenae Buszko, 1989 Elachista kosteri Traugott-Olsen, 1995 Elachista maculosella Chrétien, 1896 Elachista megagnathos Sruoga, 1990 Elachista minuta (Parenti, 2003) Elachista morandinii Huemer[verification needed] & Kaila, 2003 Elachista sicula Parenti, 1978 Elachista stenopterella Rebel, 1932 Other groups Some Elachista are divided into groups whose relationships to the two large subgenera requires further study, including:[8] "Dibrachia" group Elachista alicanta Kaila, 2005 Elachista anatoliensis Traugott-Olsen, 1990 Elachista elksourensis Kaila, 2005 Elachista kalki Parenti, 1978 (type of Dibrachia) Elachista totalbella Chrétien, 1908 "Irenicodes" group Elachista eurychora (Meyrick, 1919) (type of Irenicodes) Elachista galatheae (Viette, 1954) (type of Euproteodes) Elachista holdgatei (Bradley, 1965) Elachista hookeri (Dugdale, 1971) Elachista pumila (Dugdale, 1971) Incertae sedis Additional Elachista species are of even more uncertain relationships than usual for this genus:[9] Elachista albella[verification needed] (Chambers, 1877) (type of Hemiprosopa) Elachista altaica Sinev, 1998 Elachista baikalica Kaila, 1992 Elachista bassii Parenti, 2006 Elachista beltira Kaila, 2000 Elachista beriga Kaila, 2000 Elachista bimaculata Parenti, 1981 Elachista brevis Sruoga & J. de Prins, 2009 Elachista chelonites Meyrick, 1909 (type of Cleroptila) Elachista deresyensis Traugott-Olsen,1988 Elachista donia Kaila, 2000 Elachista gibbera Kaila, 2003 Elachista kakamegensis Sruoga & J. de Prins, 2009 Elachista kherana Kaila, 2000 Elachista longispina Sruoga & J. de Prins, 2009 Elachista maculosa Parenti, 1981 Elachista margareteae (Traugott-Olsen, 1994) Elachista mus Parenti, 1981 Elachista planca Sruoga & J. de Prins, 2009 Elachista puplesisi Sruoga, 2000 Elachista semnani Parenti, 1981 Elachista stelviella Amsel, 1932 Elachista vasrana Kaila, 2000 Elachista veruta Kaila, 2008 Synonyms Invalid scientific names (junior synonyms and others) of Elachista in the circumscription as presented here are:[10] Aphelosetia Stephens, 1834 Aphigalia Dyar, [1903] Atachia Wocke in Heinemann, [1876] Biselachista Traugott-Olsen & Schmidt Nielsen[verification needed], 1977 Cleroptila Meyrick, 1909 Cosmiotes Clemens, 1860 Cycnodia Herrich-Schäffer, 1853 Dibrachia Sinev & Sruoga, 1992 Dicasteris Meyrick, 1906 Elaschista (lapsus[verification needed]) Euproteodes Viette, 1954 Hecista Wallengren, 1881 Hemiprosopa A.F.Braun, 1948 Irenicodes Meyrick, 1919 Neaera Chambers, 1880 (non Robineau-Desvoidy, 1830: preoccupied) Paraperittia Rebel, 1916 Phigalia Chambers, 1875 (non Duponchel, 1829: preoccupied) Platyphyllis Meyrick, 1932 Ptilodoxa Meyrick, 1921 Svenssonia Parenti, 2009 Footnotes ^ AEBR (2008), FE (2009) ^ AEBR (2008), FE (2009), and see references in Savela (2008) ^ Kusdas & Reichl (1973): 194, Pitkin & Jenkins (2004), AEBR (2008), FE (2009), and see references in Savela (2008) ^ FE (2009), Wikispecies (2009-SEP-27), and see references in Savela (2008) ^ a b c d See references in Savela (2008) ^ a b c d e f g h i j k Kaila (2011) ^ a b Sorted in the presumed phylogenetic sequence (Kaila, 2011). ^ ABRS (2008), FE (2009), Wikispecies (2009-SEP-27) ^ Wikispecies (2009-SEP-27), and see references in Savela (2008) ^ AEBR (2008), FE (2009), Wikispecies (2009-SEP-27), and see references in Savela (2008) References Data related to Elachista at Wikispecies Australian Biological Resources Study (ABRS) (2008): Australian Faunal Directory – Elachista. Version of 2008-OCT-09. Retrieved 2010-MAY-01. Fauna Europaea (FE) (2009): Elachista. Version 2.1, 2009-DEC-22. Retrieved 2010-MAY-01. Kaila, Lauri (2011): Elachistine Moths of Australia (Lepidoptera: Gelechioidea, Elachistidae) (Monographs on Australian Lepidoptera Series 11). CSIRO Publishing. ISBN 978-0643103054. Preview at Google Books Kusdas, Karl & Reichl, Ernst Rudolf (eds.) (1973): Schmetterlinge Oberösterreichs (Vol. 6: Microlepidoptera) [in German]. Landwirtschaftskammer für Oberösterreich. Pitkin, Brian & Jenkins, Paul (2004): Butterflies and Moths of the World, Generic Names and their Type-species – Elachista. Version of 2004-NOV-05. Retrieved 2010-MAY-01. Savela, Markku (2008): Markku Savela's Lepidoptera and some other life forms – Elachista. Version of 2008-JUL-19. Retrieved 2010-MAY-01. Wikimedia Foundation. 2010. 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