Maranhão red-handed howler

Maranhão red-handed howler
Maranhão red-handed howler
Conservation status
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Primates
Family: Atelidae
Genus: Alouatta
Species: A. ululata
Binomial name
Alouatta ululata
Elliot, 1912
Maranhao Red-handed Howler range

Alouatta belzebul ululata

The Maranhão red-handed howler (Alouatta ululata) is an endangered species of howler monkey that is endemic to forests (for example Babaçu forests) in the northeast Brazilian states of Ceará, Maranhão and Piauí.[1] It was previously thought to be a subspecies of the Red-handed Howler, but unlike that species the Maranhão Red-handed Howler is strongly sexually dichromatic.[2]


  1. ^ a b de Oliveira, M.M. & Keirulff, M.C.M. (2008). Alouatta ululata. In: IUCN 2008. IUCN Red List of Threatened Species. Downloaded on 13 February 2009.
  2. ^ Gregorin, R. (2006). (Portuguese) Taxonomia e variação geográfica das espécies do gênero Alouatta Lacépède (Primates, Atelidae) no Brasil. Rev. Bras. Zool. 23(1).