# Leaf angle distribution

Leaf angle distribution

The Leaf Angle Distribution (or LAD) of a plant canopy refers to the mathematical description of the angular orientation of the leaves in the vegetation. Specifically, if each leaf is conceptually represented by a small flat plate, its orientation can be described with the zenith and the azimuth angles of the surface normal to that plate. If the leaf has a complex structure and is not flat, it may be necessary to approximate the actual leaf by a set of small plates, in which case there may be a number of leaf normals and associated angles. The LAD describes the statistical distribution of these angles.

Examples of Leaf Angle Distributions

Different plant canopies exhibit different LADs: For instance, grasses and willows have their leaves largely hanging vertically (such plants are said to have an erectophile LAD), while oaks tend to maintain their leaves more or less horizontally (these species are known as having a planophile LAD). In some tree species, leaves near the top of the canopy follow an erectophile LAD while those at the bottom of the canopy are more planophile. This may be interpreted as a strategy by that plant species to maximize exposure to light, an important constraint to growth and development. Yet other species (notably sunflower) are capable of reorienting their leaves throughout the day to optimize exposure to the Sun: this is known as heliotropism.

The LAD of a plant canopy has a significant impact on the reflectance, transmittance and absorption of solar light in the vegetation layer, and thus also on its growth and development. LAD can also serve as a quantitative index to monitor the state of the plants, as wilting usually results in more erectophile LADs. Models of radiation transfer need to take this distribution into account to predict, for instance, the albedo or the productivity of the canopy.

Accurately measuring the statistical properties of leaf angle distributions is not a trivial matter, especially for small leaves. Clinometers can be used but may be rather bulky or inconvenient. Most leaves are quite light and tend to move with the slightest breeze or air turbulence. Nevertheless, when these environmental conditions are suitable or can be controlled, it is possible to acquire data on leaf orientation.

This may be done, for instance, with a Spatial Coordinate Apparatus, which is an articulated mechanical device capable or recording the position in three-dimensional space of three separate points forming a small triangle. The orientation of the triangle is computed from these coordinates.

Yet another approach is to use a laser scanner: this instrument can record the angular and distance coordinates of the intersection of a laser beam with objects within its range. The LAD of canopy leaves can be derived from such measurements.

Extensive data sets of LAD have been recorded, especially during intensive field campaigns, such as the First ISLSCP Field Experiment (FIFE) or the SAFARI 2000 field campaign. See the external links below for getting access to these data.

References

* J. Ross (1981) "The Radiation Regime and Architecture of Plant Stands", 391 pp., W. Junk, Boston, Massachusetts.
* David M. Gates (1980) "Biophysical Ecology", Springer-Verlag, New York, 611 pp., ISBN 0-387-90414-X (especially pages 379-381).

* http://www.daac.ornl.gov/FIFE/Datasets/Vegetation/Leaf_Angle_Data.html
* http://www.daac.ornl.gov/S2K/guides/kt_canopy_structure.html

Wikimedia Foundation. 2010.

### Look at other dictionaries:

• Leaf — For other uses, see Leaf (disambiguation). The leaves of a Beech tree …   Wikipedia

• Hemispherical photography — Hemispherical photography, also known as fisheye or canopy photography, is a technique to estimate solar radiation and characterize plant canopy geometry using photographs taken looking upward through an extreme wide angle lens (Rich 1990).… …   Wikipedia

• Wilting — refers to the loss of rigidity of non woody parts of plants. This occurs when the turgor pressure in non lignified plant cells falls towards zero, as a result of diminished water in the cells. The process of wilting modifies the leaf angle… …   Wikipedia

• Biometeorology — is the interdisciplinary field of science that studies the interactions between the biosphere and the Earth s atmosphere on time scales of the order of seasons or shorter (by opposition to bioclimatology).Examples of relevant processesWeather… …   Wikipedia

• Glossary of environmental science — This is a glossary of environmental science.Environmental science is the study of interactions among physical, chemical, and biological components of the environment. Environmental science provides an integrated, quantitative, and… …   Wikipedia

• angiosperm — /an jee euh sperrm /, n. Bot. a plant having its seeds enclosed in an ovary; a flowering plant. Cf. gymnosperm. [ANGIO + SPERM] * * * ▪ plant Introduction       any member of the more than 300,000 species of flowering plants (division Anthophyta) …   Universalium

• Life Sciences — ▪ 2009 Introduction Zoology       In 2008 several zoological studies provided new insights into how species life history traits (such as the timing of reproduction or the length of life of adult individuals) are derived in part as responses to… …   Universalium

• climate — /kluy mit/, n. 1. the composite or generally prevailing weather conditions of a region, as temperature, air pressure, humidity, precipitation, sunshine, cloudiness, and winds, throughout the year, averaged over a series of years. 2. a region or… …   Universalium

• photography, technology of — Introduction       equipment, techniques, and processes used in the production of photographs.  The most widely used photographic process is the black and white negative–positive system (Figure 1 >). In the camera the lens projects an image of… …   Universalium

• Glossary of botanical terms — Many of the terms used in Wikipedia glossaries (often most) are already defined and explained within Wikipedia itself. However, lists like the following indicate where new articles need to be written and are also useful for looking up and… …   Wikipedia