- Precocial
In
Biology , the term precocial refers to species in which the young are relatively mature and mobile from the moment of birth or hatching. The opposite developmental strategy is called "altricial ", where the young are born helpless. Extremely precocial species may be called "superprecocial ". These three categories form a continuum, without distinct gaps between them. Precocial species are normallynidifugous , meaning that they leave the nest shortly after birth.The span between precocial and altricial species is particularly broad in
bird s. Precocial birds, including many ground-nesting species, have offspring that are born with well-ossified skeletons, with good sight, and covered with feathers. Very precocial birds can be ready to leave the nest in a short period of time following hatching (e.g. 24 hours). Many precocial chicks are not independent inthermoregulation (the ability to regulate their own body temperatures), and they depend on the attending parent(s) to brood them with body heat for a short period of time. Precocial birds find their own food, sometimes with help or instruction from the parents. Examples of precocial birds include the domesticchicken , many species ofduck s and geese, and rails andcrake s. The most extreme,superprecocial birds are themegapode s, where the newly-hatched chicks dig themselves out of the nest mound without parental assistance, and fly on the first day after hatching.Precociality is found in many other animal groups. Familiar examples of precocial
mammal s are mostungulate s, theguinea pig , and most species ofhare . This last example demonstrates that precociality is not a particularly conservative characteristic, in the evolutionary sense, since the closely relatedrabbit is highlyaltricial .Precocial species typically have a longer
gestation or incubation period than related altricial species, and smaller litters or clutches, since each offspring has to be brought to a relatively advanced (and large) state before birth or hatching.The phenomenon of imprinting studied by
Konrad Lorenz is characteristic of precocial birds.Etymology
The word "precocial" is derived from the same root as "precocious", implying in both cases early maturity. ["Webster's New World Dictionary". David B. Guralnik, Editor in Chief. New York:
Simon and Schuster , 1984.]Phylogeny
Precociality is thought to be ancestral in birds. Thus, altricial birds tend to be found in the most derived groups. There is some evidence for precociality in
Protobird s (Elzanowski, 1995.) andTroodontid s (Varricchio et al. 2002).References
Footnotes
Notations
*Starck J.M., Ricklefs R. E. Patterns of Development: The Altricial - Precocial Spectrum. In; Avian Growth and Development. Oxford University Press, New York, NY, 1998.
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