Chlaenius

Chlaenius
A: C. velutinus; B: C. spoliatus; C: C. festivus
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Family:	Carabidae
Subfamily:	Licininae
Genus:	Chlaenius Bonelli, 1810

Chlaenius is a large and diverse genus of ground beetle. It is native to the Palearctic (including Europe), the Near East, North Africa, Afrotropical region, and the Nearctic.

Species

• Chlaenius abacoides Alluaud, 1933
• Chlaenius aberanus Sternberg, 1908
• Chlaenius aberrans Bates, 1882
• Chlaenius abstersus Bates, 1873
• Chlaenius accedens Chaudoir, 1876
• Chlaenius acroxanthus Chaudoir, 1876
• Chlaenius acutecostatus Alluaud, 1933
• Chlaenius adagidensis Sternberg, 1908
• Chlaenius adametzi Kuntzen, 1913
• Chlaenius adonis Andrewes, 1923
• Chlaenius aeneocephalus Dejean, 1826
• Chlaenius aeratus Quensel, 1806
• Chlaenius aestivus Say, 1823
• Chlaenius aethiopicus Chaudoir, 1876
• Chlaenius afganus Jedlicka, 1956
• Chlaenius africanicus Kirschenhofer, 2009
• Chlaenius agiloides Jedlicka, 1935
• Chlaenius albissoni Reitter, 1908
• Chlaenius alesi Jedlicka, 1935
• Chlaenius alexanderdostali Kirschenhofer, 2008
• Chlaenius allacteus Alluaud, 1919
• Chlaenius allardi (Basilewsky, 1967)
• Chlaenius allardianus Lorenz, 1998
• Chlaenius alluaudi Fairmaire, 1901
• Chlaenius alluaudianus Jedlicka, 1957
• Chlaenius alternans (Imhoff, 1843)
• Chlaenius alternatus G. Horn, 1871
• Chlaenius alutaceus Gebler, 1829
• Chlaenius amabilis Chaudoir, 1876
• Chlaenius amarae Andrewes, 1920
• Chlaenius amauropterus Chaudoir, 1856
• Chlaenius amazonicus Chaudoir, 1876
• Chlaenius ammon (Fabricius, 1801)
• Chlaenius amoenus Dejean, 1831
• Chlaenius amplians Bates, 1891
• Chlaenius amplicollis (Basilewsky, 1953)
• Chlaenius amplipennis Chaudoir, 1876
• Chlaenius amydrus Alluaud, 1934
• Chlaenius analis (Olivier, 1795)
• Chlaenius analisimilis Pomeroy, 1932
• Chlaenius anchomenoides Bates, 1889
• Chlaenius andamanensis Andrewes, 1920
• Chlaenius androyanus Jeannel, 1949
• Chlaenius angolanus (Basilewsky, 1953)
• Chlaenius angustatus Dejean, 1831
• Chlaenius anischenkoi Kirschenhofer, 2010
• Chlaenius annulipes Bates, 1892
• Chlaenius antennatus Chaudoir, 1876
• Chlaenius anthraceus Alluaud, 1918
• Chlaenius anthracoderus Laferte-Senectere, 1851
• Chlaenius apicalis Wiedemann, 1819
• Chlaenius apollo Andrewes, 1919
• Chlaenius argentinicus Jedlicka, 1946
• Chlaenius armenus Jedlicka, 1950
• Chlaenius arnoldi Basilewsky, 1949
• Chlaenius aruwimius Bates, 1890
• Chlaenius aspericollis Bates, 1873
• Chlaenius assamensis Kirschenhofer, 2003
• Chlaenius assecla Laferte-Senectere, 1851
• Chlaenius aterrimus (Basilewsky, 1956)
• Chlaenius athleta Kryzhanovskij, 1976
• Chlaenius atratulus Mandl, 1983
• Chlaenius atratus Chaudoir, 1876
• Chlaenius atripes Chaudoir, 1876
• Chlaenius atropos Andrewes, 1941
• Chlaenius attenuatus Klug, 1833
• Chlaenius augustus Newman, 1838
• Chlaenius aurifex Basilewsky, 1949
• Chlaenius auripilis Andrewes, 1936
• Chlaenius australis Dejean, 1831
• Chlaenius azurescens Chaudoir, 1876
• Chlaenius azureulus Mandl, 1983
• Chlaenius baehri (Kirschenhofer, 1998)
• Chlaenius baeticus Rambur, 1837
• Chlaenius balthasari Jedlicka, 1935
• Chlaenius bandjermasinensis Kirschenhofer, 2008
• Chlaenius bangkokensis Kirschenhofer, 2004
• Chlaenius barkeri Csiki, 1931
• Chlaenius basilimbatus (Grundmann, 1956)
• Chlaenius baxii (Gory, 1833)
• Chlaenius beatus Bates, 1891
• Chlaenius bengalensis Chaudoir, 1856
• Chlaenius benyovszkyi (Csiki, 1931)
• Chlaenius berndjaegeri Kirschenhofer, 2008
• Chlaenius bicolor Chaudoir, 1876
• Chlaenius bifenestratus Klug, 1832
• Chlaenius bilyi Kirschenhofer, 2009
• Chlaenius bimaculatus Dejean, 1826
• Chlaenius binghami Andrewes, 1919
• Chlaenius bioculatus Chaudoir, 1856
• Chlaenius bipustulatus Boheman, 1848
• Chlaenius birmanicus Chaudoir, 1876
• Chlaenius bisignatus Dejean, 1826
• Chlaenius bivulnerus Motschulsky, 1858
• Chlaenius bocandei (Laferte-Senectere, 1852)
• Chlaenius bohemani Chaudoir, 1856
• Chlaenius boisduvalii Dejean, 1831
• Chlaenius borellyi Burgeon, 1941
• Chlaenius borgouensis Kirschenhofer, 2007
• Chlaenius bottegoi Alluaud, 1933
• Chlaenius boueti (Jeannel, 1949)
• Chlaenius boukali Kirschenhofer, 2005
• Chlaenius braminus Chaudoir, 1876
• Chlaenius brasiliensis Dejean, 1831
• Chlaenius breuningi Jedlicka, 1931
• Chlaenius brevilabris Le Conte, 1848
• Chlaenius brevior Fairmaire, 1901
• Chlaenius breviusculus Chaudoir, 1876
• Chlaenius brunnescens Kirschenhofer, 2009
• Chlaenius bulirschi Kirschenhofer, 2010
• Chlaenius buquetii (Dejean & Boisduval, 1830)
• Chlaenius buriensis Kirschenhofer, 1998
• Chlaenius caecus Dejean, 1831
• Chlaenius caeruleiceps (Bates, 1892)
• Chlaenius caeruleicollis Chaudoir, 1876
• Chlaenius caesitius Andrewes, 1923
• Chlaenius caffer Boheman, 1848
• Chlaenius callichloris Bates, 1873
• Chlaenius cambodiensis Bates, 1889
• Chlaenius camerunensis Kirschenhofer, 2003
• Chlaenius canariensis Dejean, 1831
• Chlaenius carbonatus Chaudoir, 1876
• Chlaenius caurinus (G. Horn, 1885)
• Chlaenius cavilabrum Barker, 1922
• Chlaenius cebuensis Kirschenhofer, 2008
• Chlaenius cecrops Kirschenhofer, 2009
• Chlaenius celer Chaudoir, 1876
• Chlaenius ceramensis Kirschenhofer, 2003
• Chlaenius chagga Alluaud, 1927
• Chlaenius chainatensis Kirschenhofer, 2011
• Chlaenius chalcoderus Chaudoir, 1876
• Chlaenius chalcothorax Wiedemann, 1823
• Chlaenius cham Chaudoir, 1876
• Chlaenius championi Andrewes, 1923
• Chlaenius changwatensis (Kirschenhofer, 1998)
• Chlaenius chapanus Andrewes, 1919
• Chlaenius chaudoiri G. Horn, 1876
• Chlaenius chengduensis Kirschenhofer, 2004
• Chlaenius chiangmaiensis Kirschenhofer, 2008
• Chlaenius chitwanensis Kirschenhofer, 2005
• Chlaenius chlorochrous Chaudoir, 1876
• Chlaenius chlorodius Dejean, 1826
• Chlaenius chrysocephalus P. Rossi, 1790
• Chlaenius chrysoderus Chaudoir, 1876
• Chlaenius chuji Jedlicka, 1946
• Chlaenius cinctus (Fabricius, 1781)
• Chlaenius circumdatus Brulle, 1835
• Chlaenius circumductus A. Morawitz, 1862
• Chlaenius circumscriptus Duftschmid, 1812
• Chlaenius clarksoni Barker, 1922
• Chlaenius clypeopatens (Basilewsky, 1949)
• Chlaenius coeruleus (Steven, 1809)
• Chlaenius colasi Kirschenhofer, 2008
• Chlaenius collarti Alluaud, 1933
• Chlaenius colmanti Alluaud, 1933
• Chlaenius colombensis Jedlicka, 1964
• Chlaenius columbinus Dejean, 1831
• Chlaenius comans Andrewes, 1919
• Chlaenius comes Peringuey, 1896
• Chlaenius communimacula Chaudoir in Oberthur, 1883
• Chlaenius conformis Dejean, 1831
• Chlaenius congoanus Alluaud, 1934
• Chlaenius congoensis Burgeon, 1935
• Chlaenius conradsi Kuntzen, 1913
• Chlaenius consobrinus Peringuey, 1896
• Chlaenius consors Peringuey, 1896
• Chlaenius contractus Chaudoir, 1876
• Chlaenius controversus Peringuey, 1926
• Chlaenius convexus Fairmaire, 1886
• Chlaenius coquereli Fairmaire, 1868
• Chlaenius corbetti Andrewes, 1919
• Chlaenius cordicollis Kirby, 1837
• Chlaenius cordifer Bates, 1891
• Chlaenius corrosulus Bates, 1892
• Chlaenius coscinioderus Chaudoir,
• Chlaenius coscinioderus Chaudoir, 1856
• Chlaenius cosciniophorus Chaudoir, 1876
• Chlaenius costiger Chaudoir, 1856
• Chlaenius costipennis Boheman, 1848
• Chlaenius costulatus Motschulsky, 1859
• Chlaenius coxalis Fischer Von Waldheim, 1844
• Chlaenius crebrepunctatus Chaudoir, 1856
• Chlaenius crenistriatus Chaudoir, 1876
• Chlaenius cribellatus Chaudoir, 1876
• Chlaenius cribellicollis Chaudoir, 1876
• Chlaenius cribricollis Dejean, 1831
• Chlaenius croesus (Fabricius, 1801)
• Chlaenius croyi Kirschenhofer, 2003
• Chlaenius cruciatus Dejean, 1831
• Chlaenius crudelis Peringuey, 1896
• Chlaenius cruralis Fischer Von Waldheim, 1829
• Chlaenius cubanus Chaudoir, 1876
• Chlaenius cumatilis Leconte, 1851
• Chlaenius cupreocinctus Reiche, 1847
• Chlaenius cupreolineatus Chaudoir, 1876
• Chlaenius cupreolus Fairmaire, 1901
• Chlaenius cupreopurpureus Kirschenhofer, 2009
• Chlaenius cupripennis Chaudoir, 1876
• Chlaenius cuprithorax Quedenfeldt, 1883
• Chlaenius cursor Chevrolat, 1835
• Chlaenius curtulus Kirschenhofer, 2004
• Chlaenius cyaneus Brulle, 1835
• Chlaenius cyanipennis Boheman, 1848
• Chlaenius cyanostolus Andrewes, 1924
• Chlaenius cylindricollis Dejean, 1831
• Chlaenius daer Darlington, 1968
• Chlaenius dajuensis Kirschenhofer, 2002
• Chlaenius dalibaiensis Kirschenhofer, 2008
• Chlaenius daressalaami Jedlicka, 1957
• Chlaenius darfurensis Kirschenhofer, 2007
• Chlaenius darlingensis Castelnau, 1867
• Chlaenius darlingtoni (Basilewsky, 1951)
• Chlaenius davidsoni Mandl, 1978
• Chlaenius decellei (Basilewsky, 1968)
• Chlaenius decipiens L. Dufour, 1820
• Chlaenius decorsei Alluaud, 1916
• Chlaenius dejeanii Dejean, 1831
• Chlaenius delicatus Bates, 1892
• Chlaenius deliciolus Bates, 1873
• Chlaenius delkeskampi (Basilewsky, 1956)
• Chlaenius densaticollis Fairmaire, 1901
• Chlaenius denticulatus Dejean, 1831
• Chlaenius deplanatus (Laferte-Senectere, 1851)
• Chlaenius deserticola Raffray, 1885
• Chlaenius deuvei Kirschenhofer, 2008
• Chlaenius deyrollei (Laferte-Senectere, 1851)
• Chlaenius dhawalagiriensis Kirschenhofer, 2002
• Chlaenius diabolicus (Basilewsky, 1961)
• Chlaenius dianus Jedlicka, 1957
• Chlaenius diephelus Alluaud, 1934
• Chlaenius differens Peyron, 1858
• Chlaenius difficilis Sternberg, 1908
• Chlaenius dilatatus (Motschulsky, 1864)
• Chlaenius dilutipes Reitter, 1894
• Chlaenius dimidiatus Chaudoir, 1842
• Chlaenius dinodoides Chaudoir, 1876
• Chlaenius discicollis (Laferte-Senectere, 1851)
• Chlaenius discopictus Fairmaire, 1893
• Chlaenius dispreticus (Basilewsky, 1968)
• Chlaenius dissidens Peringuey, 1926
• Chlaenius distigma Chaudoir, 1876
• Chlaenius ditulus Peringuey, 1904
• Chlaenius dives Dejean, 1826
• Chlaenius djaina Maindron, 1899
• Chlaenius dohrnii (Bertoloni, 1857)
• Chlaenius dondoensis Kirschenhofer, 2007
• Chlaenius doriae Chaudoir, 1876
• Chlaenius dorsalis Dejean, 1831
• Chlaenius dostojevskji Tschitscherine, 1895
• Chlaenius drescheri Louwerens, 1951
• Chlaenius ducalis Chaudoir, 1876
• Chlaenius dureli Maindron, 1899
• Chlaenius dusaultii (L. Dufour, 1821)
• Chlaenius duvaucelii (Dejean, 1831)
• Chlaenius ecuadoricus Jedlicka, 1946
• Chlaenius effugiens Peringuey, 1908
• Chlaenius elegans (Schmidt-Goebel, 1846)
• Chlaenius elgonensis Alluaud, 1939
• Chlaenius elisabethanus Burgeon, 1935
• Chlaenius elongatulus Kirschenhofer, 2009
• Chlaenius elongatus (Klug, 1833)
• Chlaenius emarginatus Say, 1823
• Chlaenius eneides Bates, 1892
• Chlaenius epicosmoides (Basilewsky, 1956)
• Chlaenius episcopalis Dejean, 1831
• Chlaenius epistrophus Alluaud, 1927
• Chlaenius eritreaensis Kirschenhofer, 2007
• Chlaenius erythrocnemis Chaudoir, 1876
• Chlaenius erythropus Germar, 1824
• Chlaenius eugrammus Basilewsky, 1947
• Chlaenius eurybates Bates, 1891
• Chlaenius euryscopus Bates, 1886
• Chlaenius exaratus (Basilewsky, 1949)
• Chlaenius exilis Andrewes, 1923
• Chlaenius extensus Mannerheim, 1825
• Chlaenius eyeni Basilewsky, 1949
• Chlaenius fairmairei Murray, 1858
• Chlaenius fallax (Olivier, 1795)
• Chlaenius farai Jedlicka, 1949
• Chlaenius farkaci Kirschenhofer, 2005
• Chlaenius fasciger Chaudoir in Oberthur, 1883
• Chlaenius feanus Bates, 1892
• Chlaenius femoratus Dejean, 1826
• Chlaenius fenestratus Chaudoir, 1876
• Chlaenius feronioides Murray, 1858
• Chlaenius festivus Panzer, 1796
• Chlaenius fimbriatus (Klug, 1833)
• Chlaenius finitimus Peringuey, 1908
• Chlaenius fizianus Basilewsky, 1961
• Chlaenius flavicornis Fischer Von Waldheim, 1842
• Chlaenius flaviguttatus W. S. Macleay, 1825
• Chlaenius flavipes Menetries, 1832
• Chlaenius flavofemoratus Laporte, 1834
• Chlaenius flavomaculatus (Kolbe, 1889)
• Chlaenius flavoscriptus Quedenfeldt, 1891
• Chlaenius fletcheri Andrewes, 1919
• Chlaenius formosanus Jedlicka, 1935
• Chlaenius forreri Bates, 1884
• Chlaenius frater Chaudoir, 1876
• Chlaenius fraterculus Maindron, 1899
• Chlaenius fraternus Kolbe, 1889
• Chlaenius freyellus Jedlicka, 1959
• Chlaenius freyi Jedlicka, 1960
• Chlaenius fugax Chaudoir, 1876
• Chlaenius fulgidicollis L. Dufour, 1820
• Chlaenius fulvipes (Chaudoir, 1835)
• Chlaenius fuscicornis Dejean, 1831
• Chlaenius gabonensis Kirschenhofer, 2009
• Chlaenius gabonicus Gemminger & Harold, 1868
• Chlaenius gansuensis Jedicka, 1935
• Chlaenius geayi Jeannel, 1949
• Chlaenius geisthardti Kirschenhofer, 2007
• Chlaenius gemmingeri Ballion, 1869
• Chlaenius geniculatus (Basilewsky, 1956)
• Chlaenius germanus Chaudoir, 1876
• Chlaenius gestroi Chaudoir, 1876
• Chlaenius ghesquierei Burgeon, 1935
• Chlaenius gikondo Kirschenhofer, 2009
• Chlaenius glabratus Dejean, 1831
• Chlaenius glabricollis Dejean, 1831
• Chlaenius glaucinus Kirschenhofer, 2007
• Chlaenius gonioderus Laferte-Senectere, 1851
• Chlaenius gonospilus Walker, 1871
• Chlaenius goossensi Alluaud, 1933
• Chlaenius goryi Gory, 1833
• Chlaenius gotschii Chaudoir, 1846
• Chlaenius greensladei Darlington, 1971
• Chlaenius greyanus White, 1841
• Chlaenius grundmanni (Basilewsky, 1956)
• Chlaenius guatemalenus Bates, 1882
• Chlaenius guineensis Kolbe, 1883
• Chlaenius gundlachi Chaudoir, 1876
• Chlaenius guttula Chaudoir, 1856
• Chlaenius hamatus Eschscholtz, 1829
• Chlaenius hamifer Chaudoir, 1856
• Chlaenius harpalinus Eschscholtz, 1833
• Chlaenius heidenfelderi Kirschenhofer, 2008
• Chlaenius helvetorum Burgeon, 1935
• Chlaenius hemichlorus Fairmaire, 1888
• Chlaenius henryi Andrewes, 1919
• Chlaenius herbaceus Chevrolat, 1834
• Chlaenius heyrovskyi Jedlicka, 1949
• Chlaenius hildebrandti Harold, 1880
• Chlaenius himalayicus Andrewes, 1923
• Chlaenius holomelas (Alluaud, 1915)
• Chlaenius hornburgi Kirschenhofer, 2008
• Chlaenius horni Sternberg, 1908
• Chlaenius hostilis Putzeys, 1880
• Chlaenius howa Kunckel Dherculais, 1891
• Chlaenius huedepohli Mandl, 1983
• Chlaenius humeralis Chaudoir, 1856
• Chlaenius humphreyi Alluaud, 1927
• Chlaenius hunanensis Kirschenhofer, 2003
• Chlaenius hypocrita Peringuey, 1896
• Chlaenius imitatus Reitter, 1895
• Chlaenius immaculatus Peringuey, 1885
• Chlaenius immunitus Murray, 1858
• Chlaenius imperialis Sternberg, 1908
• Chlaenius impictus Alluaud, 1933
• Chlaenius impunctifrons Say, 1823
• Chlaenius inaequalis Fairmaire, 1901
• Chlaenius inderiensis Motschulsky, 1850
• Chlaenius indicus Jedlicka, 1956
• Chlaenius indutus Klug, 1833
• Chlaenius infantulus Chaudoir, 1876
• Chlaenius infersus Peringuey, 1926
• Chlaenius inops Chaudoir, 1856
• Chlaenius insignis Chaudoir, 1876
• Chlaenius instabilis Raffray, 1886
• Chlaenius insulanus Louwerens, 1956
• Chlaenius insularis (Ueno, 1964)
• Chlaenius intermedius Chaudoir, 1856
• Chlaenius interruptus G. Horn, 1876
• Chlaenius irakensis Jedlicka, 1959
• Chlaenius iranensis Kirschenhofer, 1998
• Chlaenius itombwanus Basilewsky, 1961
• Chlaenius jactus Kirschenhofer, 2004
• Chlaenius jaechi (Kirschenhofer, 1991)
• Chlaenius jakeschi Jedlicka, 1967
• Chlaenius jamaicae Darlington, 1935
• Chlaenius janthinus L. Redtenbacher, 1844
• Chlaenius jeanneli Basilewsky, 1949
• Chlaenius jordani (Basilewsky, 1955)
• Chlaenius jucundulus (Basilewsky, 1970)
• Chlaenius junceus Andrewes, 1923
• Chlaenius jureceki Jedlicka, 1935
• Chlaenius juvencus Dejean, 1831
• Chlaenius kafakumbae Burgeon, 1935
• Chlaenius kanarae Andrewes, 1919
• Chlaenius kapangae Burgeon, 1935
• Chlaenius kashmiricus (Grundmann, 1955)
• Chlaenius kaszabi Jedlicka, 1951
• Chlaenius katanganus Burgeon, 1935
• Chlaenius keniaensis Kirschenhofer, 2008
• Chlaenius kenyerii Kirschenhofer, 2003
• Chlaenius kerkvoordeae Burgeon, 1935
• Chlaenius kilimanus (Basilewsky, 1956)
• Chlaenius kindermanni Chaudoir, 1856
• Chlaenius kira Alluaud, 1919
• Chlaenius kivuanus Basilewsky, 1949
• Chlaenius kivuensis Alluaud, 1933
• Chlaenius klapperichi Jedlicka, 1956
• Chlaenius koenigi Semenov, 1888
• Chlaenius kolariensis Maindron, 1898
• Chlaenius kolbei (Duvivier, 1892)
• Chlaenius kotys Kirschenhofer, 2004
• Chlaenius kraatzi Sternberg, 1908
• Chlaenius kryzhanovskyi Basilewsky, 1968
• Chlaenius kulti Basilewsky, 1949
• Chlaenius kurosawai Kasahara, 1986
• Chlaenius labroexcisus Basilewsky, 1948
• Chlaenius lacunosus Andrewes, 1920
• Chlaenius ladon Kirschenhofer, 1998
• Chlaenius laetiusculus Chaudoir, 1856
• Chlaenius laetoides Burgeon, 1935
• Chlaenius laetus (Fabricius, 1794)
• Chlaenius laevicollis Peringuey, 1926
• Chlaenius laevipennis Chaudoir, 1876
• Chlaenius laeviplaga Chaudoir, 1876
• Chlaenius laeviusculus Chaudoir, 1856
• Chlaenius lafertei Guerin-Meneville, 1843
• Chlaenius lamottei (Basilewsky, 1951)
• Chlaenius langsonensis Kirschenhofer, 2008
• Chlaenius laosensis Kirschenhofer, 2008
• Chlaenius laotinus Andrewes, 1919
• Chlaenius lapillus Basilewsky, 1949
• Chlaenius lastii Bates, 1886
• Chlaenius lateralis Brulle, 1838
• Chlaenius latesternalis (Basilewsky, 1956)
• Chlaenius laticollis Say, 1823
• Chlaenius latipalpis Mandl, 1992
• Chlaenius latipennis Sternberg, 1908
• Chlaenius latithorax Mannerheim, 1844
• Chlaenius lativittis Chaudoir, 1876
• Chlaenius latreillei (Laferte-Senectere, 1852)
• Chlaenius latro Laferte-Senectere, 1851
• Chlaenius lederi Reitter, 1888
• Chlaenius leishanensis Kirschenhofer, 2005
• Chlaenius leleupi (Basilewsky, 1952)
• Chlaenius leopoldi Burgeon, 1935
• Chlaenius leprieuri Gory, 1833
• Chlaenius leptopus Basilewsky, 1956
• Chlaenius leucops (Wiedemann, 1823)
• Chlaenius leucoristus Chaudoir, 1876
• Chlaenius leucoscelis Chevrolat, 1835
• Chlaenius leytensis Kirschenhofer, 2008
• Chlaenius limbatus Wiedemann, 1821
• Chlaenius limbicollis Chaudoir, 1876
• Chlaenius limbipennis Boheman, 1860
• Chlaenius lineatus Putzeys, 1880
• Chlaenius lineellus Motschulsky, 1859
• Chlaenius liocephalus Basilewsky, 1952
• Chlaenius lioderus Andrewes, 1923
• Chlaenius liothorax Alluaud, 1934
• Chlaenius lirifer Andrewes, 1941
• Chlaenius lissoderus Chaudoir, 1876
• Chlaenius lithophilus Say, 1823
• Chlaenius litongaensis Kirschenhofer, 2009
• Chlaenius loango Kirschenhofer, 2009
• Chlaenius lobozianus Alluaud, 1934
• Chlaenius loeblianus Kirschenhofer, 2004
• Chlaenius lombokensis Kirschenhofer, 2008
• Chlaenius lomii G. Muller, 1941
• Chlaenius lomsakensis Kirschenhofer, 1998
• Chlaenius longeantennatus (Basilewsky, 1951)
• Chlaenius longicornis Chaudoir, 1843
• Chlaenius longulus (Jeannel, 1949)
• Chlaenius louwerensi Andrewes, 1936
• Chlaenius loveridgei (Basilewsky, 1951)
• Chlaenius loxias Kirschenhofer, 2004
• Chlaenius lucasi Payron, 1858
• Chlaenius lucasi Peyron, 1858
• Chlaenius lucidicollis Laferte-Senectere, 1851
• Chlaenius lucidulus (Boheman, 1848)
• Chlaenius luculentus Andrewes, 1920
• Chlaenius lugens Chaudoir, 1876
• Chlaenius luisae Gestro, 1895
• Chlaenius lujai Burgeon, 1935
• Chlaenius lulengae Burgeon, 1935
• Chlaenius lunatus Dejean, 1826
• Chlaenius luteicauda Chaudoir, 1876
• Chlaenius luzonicus Chaudoir, 1856
• Chlaenius lyperus (Jeannel, 1949)
• Chlaenius macropus Chaudoir, 1876
• Chlaenius maculatus Dejean, 1826
• Chlaenius maculiceps Boheman, 1848
• Chlaenius maculiger Castelnau, 1867
• Chlaenius madecassus Csiki, 1931
• Chlaenius madrasensis Kirschenhofer, 2004
• Chlaenius makalolo Bates, 1886
• Chlaenius malachinus (Motschulsky, 1865)
• Chlaenius malcheri Emden, 1937
• Chlaenius mandli Grundmann, 1956
• Chlaenius manowianus Jedlicka, 1957
• Chlaenius manus Louwerens, 1969
• Chlaenius maowensis Kirschenhofer, 2008
• Chlaenius maputoensis Kirschenhofer, 2007
• Chlaenius marginipennis Gory, 1833
• Chlaenius marianensis Darlington, 1970
• Chlaenius marleyi Barker, 1922
• Chlaenius martinbaehri Kirschenhofer, 2008
• Chlaenius masoni Andrewes, 1923
• Chlaenius maxillosus G. Horn, 1876
• Chlaenius medioguttatus Chaudoir, 1876
• Chlaenius mediornatus (Basilewsky, 1956)
• Chlaenius melampus Menetries, 1849
• Chlaenius melancholicus Laferte-Senectere, 1851
• Chlaenius melanocnemis Basilewsky, 1949
• Chlaenius melanopterus Chaudoir, 1876
• Chlaenius melanopus Andrewes, 1923
• Chlaenius merkli Kirschenhofer, 2003
• Chlaenius meteorus Alluaud, 1939
• Chlaenius meticulosus Laferte-Senectere, 1851
• Chlaenius micans (Fabricius, 1792)
• Chlaenius microspilus (Bates, 1892)
• Chlaenius milleti Antoine, 1932
• Chlaenius milloti Jeannel, 1949
• Chlaenius moaboensis Kirschenhofer, 2009
• Chlaenius moestus Csiki, 1931
• Chlaenius moheliensis Kirschenhofer. 2009
• Chlaenius moluccensis Kirschenhofer, 2003
• Chlaenius momboensis Kirschenhofer, 2007
• Chlaenius monardi Alluaud, 1934
• Chlaenius montanus Lucas, 1859
• Chlaenius montivagus Andrewes, 1923
• Chlaenius morettoi Kirschenhofer, 2008
• Chlaenius morio Boheman, 1860
• Chlaenius morosus Laferte-Senectere, 1851
• Chlaenius mpanga Kirschenhofer, 2009
• Chlaenius muehlei Kirschenhofer, 2004
• Chlaenius multicolor Andrewes, 1919
• Chlaenius mutatus Gemminger & Harold, 1868
• Chlaenius naeviger A. Morawitz, 1862
• Chlaenius nanlingensis Deuve & Tian, 2005
• Chlaenius nanpingensis Kirschenhofer, 1999
• Chlaenius natalensis Chaudoir, 1876
• Chlaenius nebraskensis Lecont
• Chlaenius nebrioides Antoine, 1961
• Chlaenius nemoralis Say, 1823
• Chlaenius neocaledonicus Chaudoir in Oberthur, 1883
• Chlaenius neochloodes Lorenz, 1998
• Chlaenius nepalensis Hope, 1831
• Chlaenius nepos Chaudoir, 1876
• Chlaenius newdehliensis Kirschenhofer, 2002
• Chlaenius niger Randall, 1838
• Chlaenius nigerrimus Jedlicka, 1958
• Chlaenius nigrans Kirschenhofer, 2003
• Chlaenius nigratus Kirschenhofer, 2007
• Chlaenius nigricans Wiedemann, 1821
• Chlaenius nigricornis Fabricius, 1787
• Chlaenius nigripennis Chaudoir, 1856
• Chlaenius nigrita Dejean, 1826
• Chlaenius nigroscelis Chaudoir, 1856
• Chlaenius nigrosuturatus Mandl, 1978
• Chlaenius nilgiricus Andrewes, 1919
• Chlaenius nimbanus (Basilewsky, 1951)
• Chlaenius nitidiceps Dejean, 1826
• Chlaenius nitidicollis Dejean, 1826
• Chlaenius nitidifrons Fairmaire, 1901
• Chlaenius nitidulus Schrank, 1781
• Chlaenius noeli Alluaud, 1933
• Chlaenius noguchii Bates, 1873
• Chlaenius notabilis Laferte-Senectere, 1851
• Chlaenius nubicus Chaudoir, 1876
• Chlaenius nyassaensis Jedlicka, 1957
• Chlaenius nyika Kirschenhofer, 2009
• Chlaenius obenbergeri Jedlicka, 1935
• Chlaenius oberthuri Sternberg, 1908
• Chlaenius obesus Laferte-Senectere, 1851
• Chlaenius obliquatus Barker, 1922
• Chlaenius oblongus Dejean, 1826
• Chlaenius obscuripennis Chevrolat, 1835
• Chlaenius obscurus Klug, 1832
• Chlaenius obsidianoides Alluaud, 1935
• Chlaenius obsidianus Alluaud, 1935
• Chlaenius obtusus Dejean, 1831
• Chlaenius occultus Sloane, 1907
• Chlaenius ochroperas Bates, 1892
• Chlaenius ocreatus Bates, 1873
• Chlaenius oculatus (Fabricius, 1801)
• Chlaenius oligochrysus Alluaud, 1934
• Chlaenius olivieri Crotch, 1871
• Chlaenius olthofi Darlington, 1968
• Chlaenius omochlorus Andrewes, 1931
• Chlaenius oneili Barker, 1922
• Chlaenius oodioides Chaudoir, 1876
• Chlaenius opacipennis Chaudoir, 1876
• Chlaenius ophonoides Fairmaire, 1843
• Chlaenius orbicollis Chaudoir, 1876
• Chlaenius orbiculicollis Barker, 1922
• Chlaenius orbus G. Horn, 1871
• Chlaenius orphanus Peringuey, 1908
• Chlaenius ostrinus Andrewes, 1924
• Chlaenius ovalipennis Quedenfeldt, 1883
• Chlaenius ovampo Peringuey, 1892
• Chlaenius overlaeti Burgeon, 1935
• Chlaenius pacholatkoi Kirschenhofer, 2003
• Chlaenius pachys Chaudoir, 1876
• Chlaenius paenulatus Erichson, 1843
• Chlaenius pallidicornis Ballion, 1871
• Chlaenius pallipes (Gebler, 1823)
• Chlaenius palpalis Laferte-Senectere, 1851
• Chlaenius paluensis Kirschenhofer, 2011
• Chlaenius pan Darlington, 1968
• Chlaenius panagaeoides Chaudoir, 1876
• Chlaenius panjabensis Kirschenhofer, 1998
• Chlaenius parallelus Dejean, 1831
• Chlaenius parepus Jedlicka, 1957
• Chlaenius paromius Basilewsky, 1949
• Chlaenius patricius Harold, 1879
• Chlaenius patrizii (Basilewsky, 1953)
• Chlaenius patruelis Leconte, 1844
• Chlaenius pauxillus Kirschenhofer, 2002
• Chlaenius paykulli Crotch, 1870
• Chlaenius pectinipes Bates, 1892
• Chlaenius peltastes Jedlicka, 1935
• Chlaenius pennsylvanicus Say, 1823
• Chlaenius pericallus L. Redtenbacher, 1867
• Chlaenius peringueyi Kuntzen, 1919
• Chlaenius perpunctatus Kuntzen, 1913
• Chlaenius perrieri (Fairmaire, 1899)
• Chlaenius persimilis Chaudoir, 1876
• Chlaenius perspicillaris Erichson, 1843
• Chlaenius peruanus Erichson, 1847
• Chlaenius peterseni (Louwerens, 1967)
• Chlaenius phaenoderus Chaudoir, 1876
• Chlaenius pharaonis Motschulsky, 1865
• Chlaenius phenax Basilewsky, 1949
• Chlaenius philemon Andrewes, 1936
• Chlaenius piceus Chaudoir, 1876
• Chlaenius pictus Chaudoir, 1856
• Chlaenius piligenys Liu & Liang, 2010
• Chlaenius pilosicoerulans Mandl, 1989
• Chlaenius planipennis Chaudoir, 1876
• Chlaenius planulatus Bates, 1884
• Chlaenius platensis G. R. Waterhouse, 1841
• Chlaenius platyderus Chaudoir, 1856
• Chlaenius platynoides Alluaud, 1934
• Chlaenius plausibilis Basilewsky, 1949
• Chlaenius pleuroderus Chaudoir in Oberthur, 1883
• Chlaenius poecilinus Bates, 1892
• Chlaenius porinus Alluaud, 1929
• Chlaenius porphyrius Bates, 1891
• Chlaenius porrectus Chaudoir, 1876
• Chlaenius posticalis Motschulsky, 1854
• Chlaenius posticus (Fabricius, 1798)
• Chlaenius postmaculatus (Basilewsky, 1952)
• Chlaenius postscriptus Bates, 1873
• Chlaenius pradieri Chaudoir, 1876
• Chlaenius praefectus Bates, 1873
• Chlaenius prasinus Dejean, 1826
• Chlaenius pratensis Chaudoir, 1876
• Chlaenius pretiosus Chaudoir, 1856
• Chlaenius principalis Sternberg, 1908
• Chlaenius privatus Bates, 1892
• Chlaenius privatus Bates, 1892
• Chlaenius probsti Kirschenhofer, 2004
• Chlaenius prolixus Erichson, 1843
• Chlaenius propinquus Csiki, 1931
• Chlaenius prostenus Bates, 1873
• Chlaenius prostenus Bates, 1873
• Chlaenius protensus Chaudoir, 1876
• Chlaenius pseudocruciatus Kirschenhofer, 2009
• Chlaenius pseudocupreolus Kirschenhofer, 2010
• Chlaenius pseudoglabratus Kirschenhofer, 2009
• Chlaenius pseudolulengae Kirschenhofer, 2009
• Chlaenius pseudopalpalis Basilewsky, 1949
• Chlaenius pseudoraffrayi Basilewsky, 1949
• Chlaenius pterostichoides Andrewes, 1941
• Chlaenius ptuchodes Andrewes, 1923
• Chlaenius puberulus Boheman, 1848
• Chlaenius pubifer Chaudoir, 1876
• Chlaenius pubipennis Chaudoir, 1856
• Chlaenius puchneri Kirschenhofer, 2007
• Chlaenius pudicus (Fabricius, 1801)
• Chlaenius pugni Camerano, 1879
• Chlaenius pulchellus Boheman, 1848
• Chlaenius pulcher Nietner, 1857
• Chlaenius pulchriceps Fairmaire, 1901
• Chlaenius pumilio Kolbe, 1889
• Chlaenius punctatostriatus Chaudoir, 1856
• Chlaenius puncticollis Dejean, 1826
• Chlaenius purpureus Chaudoir, 1876
• Chlaenius purpuricollis Randall, 1838
• Chlaenius pusillus Say, 1823
• Chlaenius putzeysi Chaudoir, 1876
• Chlaenius pyrrhopodus Fairmaire, 1903
• Chlaenius pyrrhos Kirschenhofer, 2004
• Chlaenius quadricolor (Olivier, 1790)
• Chlaenius quadrimaculatus (Boheman, 1848)
• Chlaenius quadrinotatus Dejean, 1826
• Chlaenius quadriornatus Basilewsky, 1956
• Chlaenius quadrisignatus Boheman, 1860
• Chlaenius quadrisulcatus Paykull, 1790
• Chlaenius quadrivittatus Jedlicka, 1958
• Chlaenius radama Kunckel, 1891
• Chlaenius raffrayi Chaudoir, 1876
• Chlaenius rambouseki Lutshnik, 1933
• Chlaenius ranavalonae Csiki, 1931
• Chlaenius rebellis Peringuey, 1926
• Chlaenius reichei (Laferte-Senectere, 1851)
• Chlaenius reitteri Jakobson, 1906
• Chlaenius retropictus Fairmaire, 1901
• Chlaenius rhodesianus Peringuey, 1899
• Chlaenius richardsi Ali, 1967
• Chlaenius riparius Lorenz, 1998
• Chlaenius ripicola Andrewes, 1937
• Chlaenius robustus Boheman, 1848
• Chlaenius rodriguezi Chaudoir, 1876
• Chlaenius roeschkei Sternberg, 1908
• Chlaenius rotundatulus Lorenz, 1998
• Chlaenius rotundithorax Liu & Kavanaugh, 2010
• Chlaenius rotundus Andrewes, 1920
• Chlaenius rubenticollis Kirschenhofer, 2009
• Chlaenius rubricrus Alluaud, 1916
• Chlaenius rudesculptus Chaudoir, 1876
• Chlaenius rudicollis Chaudoir, 1876
• Chlaenius ruficauda Chaudoir, 1856
• Chlaenius rufifemoratus Macleay, 1825
• Chlaenius rufithorax Wiedemann, 1821
• Chlaenius rufomarginatus Dejean, 1831
• Chlaenius rugulosus Nietner, 1857
• Chlaenius rukwaensis Kirschenhofer, 2007
• Chlaenius ruvumaensis Kirschenhofer, 2008
• Chlaenius rysonotus Fairmaire, 1901
• Chlaenius sabahensis Kirschenhofer, 1998
• Chlaenius saginatus Laferte-Senectere, 1851
• Chlaenius sagittarius Dejean, 1831
• Chlaenius salisburiensis Barker, 1922
• Chlaenius sallei Chaudoir, 1876
• Chlaenius samoensis Csiki, 1915
• Chlaenius sangaicus Alluaud, 1929
• Chlaenius sankuruensis Burgeon, 1935
• Chlaenius savanicola (Basilewsky, 1968)
• Chlaenius scabricollis Chevrolat, 1833
• Chlaenius scapularis Chaudoir, 1876
• Chlaenius scheerpeltzi (Basilewsky, 1956)
• Chlaenius schillhammeri Kirschenhofer, 2003
• Chlaenius schmidti Gestro, 1895
• Chlaenius schoenherri (Dejean, 1831)
• Chlaenius schoutedeni Burgeon, 1935
• Chlaenius schuelei Kirschenhofer, 2007
• Chlaenius sciakyi Kirschenhofer, 2004
• Chlaenius scotti Alluaud, 1937
• Chlaenius sculptilis Bates, 1886
• Chlaenius scutellaris Harold, 1880
• Chlaenius sehnali Kirschenhofer, 2008
• Chlaenius seiferti Kirschenhofer, 2005
• Chlaenius sellatus Dejean, 1831
• Chlaenius semenowi Tschitscherine, 1895
• Chlaenius semicupreus Basilewsky, 1949
• Chlaenius semicyaneus Solsky, 1874
• Chlaenius semipurpureus Motschulsky, 1865
• Chlaenius senegalensis Dejean, 1831
• Chlaenius sennaariensis Chaudoir, 1856
• Chlaenius sericeus (Forster, 1771)
• Chlaenius sericimicans Chaudoir, 1876
• Chlaenius sexguttatus Andrewes, 1919
• Chlaenius sexmaculatus Dejean, 1831
• Chlaenius seyrigi Alluaud, 1935
• Chlaenius siccus Darlington, 1968
• Chlaenius sichuanensis Kirschenhofer, 2008
• Chlaenius sierraleonensis Kirschenhofer, 2008
• Chlaenius siffointei (Basilewsky, 1968)
• Chlaenius silvestrii G. Muller, 1942
• Chlaenius simba Alluaud, 1929
• Chlaenius simbabwensis Kirschenhofer, 2008
• Chlaenius similatus Boheman, 1848
• Chlaenius similis Chaudoir, 1856
• Chlaenius simplex Wiedemann, 1821
• Chlaenius simulatorius Barker, 1922
• Chlaenius sinensis Chaudoir, 1856
• Chlaenius sivorii Chaudoir, 1876
• Chlaenius sobrinus Dejean, 1826
• Chlaenius soccatus Say, 1830
• Chlaenius sokotranus Csiki, 1931
• Chlaenius solitarius Say, 1823
• Chlaenius sollicitus Laferte-Senectere, 1851
• Chlaenius somaliae Basilewsky, 1956
• Chlaenius somereni Alluaud, 1929
• Chlaenius soricinus Gerstaecker, 1867
• Chlaenius sparsepunctatus Chaudoir, 1876
• Chlaenius spathulifer (Bates, 1873)
• Chlaenius specularis Emden, 1937
• Chlaenius splendidus Dejean, 1831
• Chlaenius spoliatus P. Rossi, 1792
• Chlaenius startellus Basilewsky, 1949
• Chlaenius steveni Quensel, 1806
• Chlaenius stichai Jedlicka, 1949
• Chlaenius straneoi Basilewsky, 1949
• Chlaenius stschukini Menetries, 1837
• Chlaenius stungtrengensis Kirschenhofer, 2004
• Chlaenius stygius (Laferte-Senectere, 1851)
• Chlaenius suavis Alluaud, 1939
• Chlaenius subcostatus W. J. Macleay, 1864
• Chlaenius suberbiei (Jeannel, 1949)
• Chlaenius subferrugineus (Kirschenhofer, 2003)
• Chlaenius subovatus Chaudoir, 1876
• Chlaenius subsulcatus Dejean, 1831
• Chlaenius suensoni Mandl, 1992
• Chlaenius sulcicollis (Paykull, 1798)
• Chlaenius sulcipennis Dejean, 1826
• Chlaenius sumptuosus Alluaud, 1934
• Chlaenius superbus Sternberg, 1908
• Chlaenius superstes Peringuey, 1926
• Chlaenius suppletor Bates, 1891
• Chlaenius surdipennis (Jeannel, 1949)
• Chlaenius suvorovi Semenov, 1912
• Chlaenius swahilius Bates, 1886
• Chlaenius syangyaensis Kirschenhofer, 2004
• Chlaenius sykesi Hope, 1835
• Chlaenius synaptus Alluaud, 1918
• Chlaenius syriacus Chaudoir, 1876
• Chlaenius systolocranioides Alluaud, 1933
• Chlaenius tamdaoensis Kirschenhofer, 2003
• Chlaenius tanahrataensis Kirschenhofer, 2008
• Chlaenius teani Gestro, 1881
• Chlaenius tecospilus Alluaud, 1933
• Chlaenius tenellus Klug, 1832
• Chlaenius tenuicollis (Fabricius, 1801)
• Chlaenius tenuilimbatus Ballion, 1871
• Chlaenius tenuis Fairmaire, 1901
• Chlaenius terminatus Dejean, 1826
• Chlaenius terminatus Dejean, 1826
• Chlaenius testaceicrus Fairmaire, 1891
• Chlaenius tetracelis Alluaud, 1933
• Chlaenius tetragonoderus Chaudoir, 1876
• Chlaenius tetraphacus Alluaud, 1918
• Chlaenius tetraspilus Alluaud, 1918
• Chlaenius thieleni (Kirschenhofer, 1998)
• Chlaenius thiesensis Kirschenhofer, 2007
• Chlaenius tibialis Dejean, 1826
• Chlaenius timorensis Darlington, 1971
• Chlaenius tinantae Burgeon, 1935
• Chlaenius titschacki Jedlicka, 1946
• Chlaenius tomentosus Say, 1823
• Chlaenius toubaensis Kirschenhofer, 2008
• Chlaenius touzalini Andrewes, 1920
• Chlaenius trachys Andrewes, 1923
• Chlaenius transfuga Chaudoir, 1876
• Chlaenius transversalis Dejean, 1831
• Chlaenius trapezicollis Chaudoir, 1856
• Chlaenius treichi Alluaud, 1916
• Chlaenius trichrous Alluaud, 1918
• Chlaenius tricolor Dejean, 1826
• Chlaenius trigonotomoides Emden, 1928
• Chlaenius trinotatus Laferte-Sinectere, 1851
• Chlaenius tripustulatus (Dejean, 1831)
• Chlaenius tristis Schaller, 1783
• Chlaenius tschitscherini Jedlicka, 1952
• Chlaenius tshibindensis Burgeon, 1935
• Chlaenius tudicus Andrewes, 1919
• Chlaenius tuky Basilewsky, 1949
• Chlaenius uluguruanus Basilewsky, 1956
• Chlaenius umtalianus (Peringuey, 1904)
• Chlaenius unicolor Chaudoir, 1856
• Chlaenius uninotatus Andrewes, 1919
• Chlaenius unyorensis Alluaud, 1934
• Chlaenius uzungwensis Basilewsky, 1951
• Chlaenius vadoni (Basilewsky, 1950)
• Chlaenius validicornis Boheman, 1848
• Chlaenius varians Chaudoir, 1876
• Chlaenius variicornis A. Morawitz, 1863
• Chlaenius variipes Chaudoir, 1856
• Chlaenius variolosus Kirschenhofer, 2008
• Chlaenius vartianorum Mandl, 1989
• Chlaenius velocipes Chaudoir, 1876
• Chlaenius venustulus Dejean, 1831
• Chlaenius vertagoides (Laferte-Senectere, 1851)
• Chlaenius veselyi Kirschenhofer, 2004
• Chlaenius vestitus (Paykull, 1790)
• Chlaenius vethi Bates, 1889
• Chlaenius viangchanensis Kirschenhofer, 1998
• Chlaenius victor Andrewes, 1928
• Chlaenius victoriae Kirschenhofer, 2009
• Chlaenius viduus G. Horn, 1871
• Chlaenius vietnamensis Kirschenhofer, 2008
• Chlaenius vietnami Jedlicka, 1966
• Chlaenius violaceipennis Chaudoir, 1876
• Chlaenius violatus Gemminger & Harold, 1868
• Chlaenius virens Rambur, 1837
• Chlaenius virescens Chaudoir, 1835
• Chlaenius virgulatus (Jeannel, 1949)
• Chlaenius virgulifer Chaudoir, 1876
• Chlaenius viridicollis Reiche, 1843
• Chlaenius viridis Menetries, 1832
• Chlaenius vitalisi Andrewes, 1919
• Chlaenius vitticollis Boheman, 1848
• Chlaenius vividus Chaudoir, 1876
• Chlaenius vulneratus Dejean, 1831
• Chlaenius waddellii Murray, 1858
• Chlaenius wallacei Chaudoir, 1876
• Chlaenius wegneri Louwerens, 1953
• Chlaenius wenchuanensis Kirschenhofer, 2008
• Chlaenius wittei Burgeon, 1937
• Chlaenius wittmerianus Mandl, 1978
• Chlaenius wrasei (Kirschenhofer, 1997)
• Chlaenius xanthacrus Wiedemann, 1823
• Chlaenius xanthomerus Alluaud, 1918
• Chlaenius xanthospilus (Wiedemann, 1821)
• Chlaenius yangonensis Kirschenhofer, 2005
• Chlaenius yunnanulus Mandl, 1992
• Chlaenius zanzibaricus Chaudoir in Oberthur, 1883
• Chlaenius ziloensis Basilewsky, 1970
• Chlaenius zygogrammus Laferte-Senectere, 1851

References

"Chlaenius Bonelli, 1810". Carabidae of the World. 2011. http://carabidae.pro/carabidae/chlaenius-bonelli-1810.html. Retrieved 13 Jul 2011. 

External links

• Chlaenius at Fauna Europaea
• Chlaenius at BugGuide (North America)