G protein-coupled receptor 4, also known as GPR4, is a human gene.cite web | title = Entrez Gene: GPR4 G protein-coupled receptor 4| url =| accessdate = ]

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*cite journal | author=An S, Tsai C, Goetzl EJ |title=Cloning, sequencing and tissue distribution of two related G protein-coupled receptor candidates expressed prominently in human lung tissue. |journal=FEBS Lett. |volume=375 |issue= 1-2 |pages= 121–4 |year= 1996 |pmid= 7498459 |doi=
*cite journal | author=Heiber M, Docherty JM, Shah G, "et al." |title=Isolation of three novel human genes encoding G protein-coupled receptors. |journal=DNA Cell Biol. |volume=14 |issue= 1 |pages= 25–35 |year= 1995 |pmid= 7832990 |doi=
*cite journal | author=Mahadevan MS, Baird S, Bailly JE, "et al." |title=Isolation of a novel G protein-coupled receptor (GPR4) localized to chromosome 19q13.3. |journal=Genomics |volume=30 |issue= 1 |pages= 84–8 |year= 1996 |pmid= 8595909 |doi= 10.1006/geno.1995.0013
*cite journal | author=Zhu K, Baudhuin LM, Hong G, "et al." |title=Sphingosylphosphorylcholine and lysophosphatidylcholine are ligands for the G protein-coupled receptor GPR4. |journal=J. Biol. Chem. |volume=276 |issue= 44 |pages= 41325–35 |year= 2001 |pmid= 11535583 |doi= 10.1074/jbc.M008057200
*cite journal | author=Strausberg RL, Feingold EA, Grouse LH, "et al." |title=Generation and initial analysis of more than 15,000 full-length human and mouse cDNA sequences. |journal=Proc. Natl. Acad. Sci. U.S.A. |volume=99 |issue= 26 |pages= 16899–903 |year= 2003 |pmid= 12477932 |doi= 10.1073/pnas.242603899
*cite journal | author=Lum H, Qiao J, Walter RJ, "et al." |title=Inflammatory stress increases receptor for lysophosphatidylcholine in human microvascular endothelial cells. |journal=Am. J. Physiol. Heart Circ. Physiol. |volume=285 |issue= 4 |pages= H1786–9 |year= 2003 |pmid= 12805023 |doi= 10.1152/ajpheart.00359.2003
*cite journal | author=Ludwig MG, Vanek M, Guerini D, "et al." |title=Proton-sensing G-protein-coupled receptors. |journal=Nature |volume=425 |issue= 6953 |pages= 93–8 |year= 2003 |pmid= 12955148 |doi= 10.1038/nature01905
*cite journal | author=Bektas M, Barak LS, Jolly PS, "et al." |title=The G protein-coupled receptor GPR4 suppresses ERK activation in a ligand-independent manner. |journal=Biochemistry |volume=42 |issue= 42 |pages= 12181–91 |year= 2003 |pmid= 14567679 |doi= 10.1021/bi035051y
*cite journal | author=Sin WC, Zhang Y, Zhong W, "et al." |title=G protein-coupled receptors GPR4 and TDAG8 are oncogenic and overexpressed in human cancers. |journal=Oncogene |volume=23 |issue= 37 |pages= 6299–303 |year= 2004 |pmid= 15221007 |doi= 10.1038/sj.onc.1207838
*cite journal | author=Gerhard DS, Wagner L, Feingold EA, "et al." |title=The status, quality, and expansion of the NIH full-length cDNA project: the Mammalian Gene Collection (MGC). |journal=Genome Res. |volume=14 |issue= 10B |pages= 2121–7 |year= 2004 |pmid= 15489334 |doi= 10.1101/gr.2596504
*cite journal | author=Kim KS, Ren J, Jiang Y, "et al." |title=GPR4 plays a critical role in endothelial cell function and mediates the effects of sphingosylphosphorylcholine. |journal=FASEB J. |volume=19 |issue= 7 |pages= 819–21 |year= 2005 |pmid= 15857892 |doi= 10.1096/fj.04-2988fje
*cite journal | author=Qiao J, Huang F, Naikawadi RP, "et al." |title=Lysophosphatidylcholine impairs endothelial barrier function through the G protein-coupled receptor GPR4. |journal=Am. J. Physiol. Lung Cell Mol. Physiol. |volume=291 |issue= 1 |pages= L91–101 |year= 2006 |pmid= 16461426 |doi= 10.1152/ajplung.00508.2005
*cite journal | author=Huang F, Mehta D, Predescu S, "et al." |title=A novel lysophospholipid- and pH-sensitive receptor, GPR4, in brain endothelial cells regulates monocyte transmigration. |journal=Endothelium |volume=14 |issue= 1 |pages= 25–34 |year= 2007 |pmid= 17364894 |doi= 10.1080/10623320601177288
*cite journal | author=Zou Y, Kim CH, Chung JH, "et al." |title=Upregulation of endothelial adhesion molecules by lysophosphatidylcholine. Involvement of G protein-coupled receptor GPR4. |journal=FEBS J. |volume=274 |issue= 10 |pages= 2573–84 |year= 2007 |pmid= 17437524 |doi= 10.1111/j.1742-4658.2007.05792.x
*cite journal | author=Tobo M, Tomura H, Mogi C, "et al." |title=Previously postulated "ligand-independent" signaling of GPR4 is mediated through proton-sensing mechanisms. |journal=Cell. Signal. |volume=19 |issue= 8 |pages= 1745–53 |year= 2007 |pmid= 17462861 |doi= 10.1016/j.cellsig.2007.03.009

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