Alchornea glandulosa

Alchornea glandulosa

Taxobox
name = "Alchornea glandulosa"



image_width = 200px
status = NE
status_system = iucn3.1
regnum = Plantae
divisio = Magnoliophyta
classis = Magnoliopsida
ordo = Malpighiales
familia = Euphorbiaceae
subfamilia = Acalyphoideae
tribus = Alchorneae
subtribus = Alchorneinae
genus = "Alchornea"
species = "A. glandulosa"
binomial = "Alchornea glandulosa"
binomial_authority = Poepp. & Endl.
synonyms = "Alchornea irucurama" Casar.
"Alchornea pittieri" Pax
"Alchornea subrotunda" Baill.
"Alchornea umboensis" Croizat

"Alchornea glandulosa" is a tree species of the Acalyphoideae native to South America, growing for example in southern Brazil from Minas Gerais to Rio Grande do Sul. It is locally known as "tamanqueiro", "tapiá" or "amor seco". This gnarled tree grows preferentially in riparian forest, where it a common pioneer species growing to a heigth a height of 10-20 m. It is essentially evergreen, though in the hot austral summer months there is a more pronounced changeover of leaves, and branches are denuded to some extent. [Conegero "et al." (2003), Pascotto "et al." (2006)]

The fruit is about 8.7 mm long by 5.9 mm wide on average, and contains one round seed measuring about 4.45 mm in diameter; very rarely a second seed develops. This sticks out of an aril at the fruit's tip; when ripe, the seedcoat turns bright red and the fruit somewhat resembles that of a yew with a larger and more prominent seed. Fruit ripen in the summer months, roughly between September/October and December/January in S Brazil, and as the trees bear less leaves at that time than otherwise, the bright red fruit are easily spotted.Pascotto "et al." (2006)]

This tree is often cut down for timber, but it is also useful as a honey plant. Also, its leaves contain compounds of medical interest. In folk medicine, "Alchornea" species are used to treat assorted skin diseases, diarrhea, inflammations, leprosy and rheuma. Scientific studies have confirmed most of these effects, and also found extracts of certain species to kill off trypanosoma, some bacteria and fungi, and cancer cells; the latter properties have also been tested in "A. glandulosa". Compounds of interest in "A. glandulosa" include the phytosterols β-sitosterol and stigmasterol, the terpenoid loliolide, the guanidine alkaloid N-1,N-2,N-3-triisopentenylguanidine, and the phenolic compound corilagin. [Conegero "et al." (2003)]

Given the fruit's attractive color and the conspicuous display at the branch-tips, the this tree appears to be distributed by birds which eat the fruit and spread the seeds. Perching birds, namely tanagers (Thraupidae), thrushes (Turdidae) and tyrant flycatchers (Tyrannidae), are most commonly seen to feed on the fruit. Some birds, such as the Silver-beaked Tanager ("Ramphocelus carbo"), prune off the seedcoat and eat it, discarding the seed, but most swallow the entire fruit. Species such as the Swallow Tanager ("Tersina viridis") and the Pale-breasted Thrush ("Turdus leucomelas"), but perhaps most of all the Sayaca Tanager ("Thraupis sayaca"), appear to be particularly fond of them. Indeed, the Sayaca Tanager will defend richly fruiting "A. glandulosa" trees against similar-sized birds such as the Blue Dacnis ("Dacnis cayana") or the Red-eyed Vireo ("Vireo olivaceus"). It may be that at least locally, "T. sayaca" is crucial for the tree's reproduction and survival.

"A. glandulosa" fruit are also significant food of certain migrant birds in their winter quarters. In particular the Red-eyed Vireo and the White-necked Thrush ("Turdus albicollis") have been noted to be fond of them, and Swainson's Flycatchers ("Myiarchus swainsoni") visit the trees very often too. But as it does not relish the fruits very much, the latter species is perhaps more attracted to insects living on the tree.

ee also

*List of honey plants

Footnotes

References

* (2003): Constituintes químicos de "Alchornea glandulosa" (Euphorbiaceae) [Chemical contituents of "Alchornea glandulosa" (Euphorbiaceae)] . "Quím. Nova" 26(6): 825-827 [Portuguese with English abstract] . doi|10.1590/S0100-40422003000600008 [http://www.scielo.br/pdf/qn/v26n6/a08v26n6.pdf PDF fulltext]
* (2006): Avifauna dispersora de sementes de "Alchornea glandulosa" (Euphorbiaceae) em uma área de mata ciliar no estado de São Paulo [Seed dispersal of "Alchornea glandulosa" (Euphorbiaceae) by birds in a gallery forest in São Paulo, southeastern Brazil.] . "Revista Brasileira de Ornitologia" 14(3): 291-296 [Portuguese with English abstract] . [http://www.ararajuba.org.br/sbo/ararajuba/artigos/Volume143/ara143not5.pdf PDF fulltext]


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