name = Ratardinae
subfamilia = Ratardinae
subdivision_ranks = Genera
Ratarda" Moore, 1879
**="Shisa" Strand, 1917
Callosiope" Hering, 1925
Sumatratarda" Kobes and Ronkay, 1990
Taxonomy and systematics
Ratardinae is a small subfamily of
moths formerly placed in its own family Ratardidae and related to (and often included within) Cossidae. There are three genera, one quite recently described (Kobes and Ronkay, 1990). One species, "Shisa" excellens" was originally placed in Lymantriidae(Owada, 1993; Holloway, 1998: 9). The moths are large with rounded wings and strongly spotted wing patterns, and "pectinate" antennae. The relationships of this group to other Cossoideaneeds reassessment, once suitable samples are available, with moleculardata [http://www.leptree.net/statusmatrix/showall.php] .
There are about thirteen, relictually distributed species restricted to
Southeast Asia, occurring in Borneo, Sumatra, Peninsular Malaysia, northeastern Himalayasand Taiwan(Holloway, 1986: 41-42; Heppner and Wang, 1987; Heynderycx, 2003).
The adults are very rarely found, feeble- flying and occasionally are attracted to light but more likely to be found flying by day (Holloway, 1986: 41-42).
The biology is unknown (Edwards et al. 1999: 193-194) but it has been suggested based on female morphology that like the probably related
Metarbelinaethey might feed in bark (Holloway et al., 1986: 42).
These large moths are so incredibly rarely found and their habitats under such massive threat from large-scale conversion of rainforest in South-East Asia that their conservation status should be seriously considered and dedicated surveys conducted to assess their distribution and biology. One species ("Ratarda melanoxantha") is probably protected by virtue of its occurrence in
Mount KinabaluNational Park on Borneowhere it was found once (Holloway, 1986) .
* [http://web.archive.org/web/20060116092931/http://www.mothsofborneo.com/part-1/ratardidae/ratardidae.php The Moths of Borneo]
* [http://internt.nhm.ac.uk/jdsml/research-curation/projects/lepindex/list.dsml?searchPageURL=index.dsml&UserID=&UserName=&sort=SCIENTIFIC_NAME_on_card&Current_superfamily=Cossoidea&Current_family=Cossidae&Current_subfamily=Ratardinae&Current_tribe=&recLimit=30 NHM Lepindex]
* [http://www.funet.fi/pub/sci/bio/life//insecta/lepidoptera/ditrysia/cossoidea/cossidae/ratardinae/index.html Markku Savela's website]
*Edwards, E.D., Gentili, P., Horak, M., Kristensen, N.P. and Nielsen, E.S. (1999). The cossoid/sesioid assemblage. Ch. 11, pp. 181-195 in Kristensen, N.P. (Ed.). "Lepidoptera, Moths and Butterflies". Volume 1: Evolution, Systematics, and Biogeography. Handbuch der Zoologie. Eine Naturgeschichte der Stämme des Tierreiches / Handbook of Zoology. A Natural History of the phyla of the Animal Kingdom. Band / Volume IV Arthropoda: Insecta Teilband / Part 35: 491 pp. Walter de Gruyter, Berlin, New York.
*Heynderycx, J. 2003. Les Ratardinae. "Lambillionea;;, 103(1): 133-134.
*Heppner J.B. and Wang, H.Y. 1987. A rare moth, "Ratarda tertia" Strand (Lepidoptera: Ratardidae), from Palin Taiwan. "Táiwa-n she(nglì bówùgua(n bànniánka-n (Táiwa-n she(nglì bówùgua(n bànniánka-n)", 40: 91-94.
*Holloway, J.D. (1986). "The Moths of Borneo: Key to Families: Families Cossiae, Metarbelidae, Ratardidae, Dudgeonidae, Epipyropidae and Limacodidae". "Malayan Nauture Journal", 40: 1-166.
*Holloway, J.D. (1998). "The Moths of Borneo: Families Castniidae, Callidulidae, Drepanidae and Uraniidae". "Malayan Nauture Journal", 52: 1-155.
*Kobes, L.W.R. and Ronkay, L. (1990). "The Ratardidae of Sumatra", "Heterocera Sumatrana", 6: 79-100.
*Owada, M. (1993). The systematic position of "Shisa excellens" (Lepidoptera and Ratardidae). "Japanese Journal of Entomology", 61(2): 251-260.
Provisional list of species (based on Lepindex)
Callosiope banghaasi" Hering, 1925
Ratarda flavimargo" Hering, 1925 [http://images.google.co.uk/imgres?imgurl=http://www.mothsofborneo.com/part-1/images/plate4/11.jpg&imgrefurl=http://www.mothsofborneo.com/part-1/ratardidae/ratardidae.php&h=119&w=201&sz=17&hl=en&start=1&tbnid=elOSfcoRk9my5M:&tbnh=62&tbnw=104&prev=/images%3Fq%3DRatarda%26gbv%3D2%26svnum%3D10%26hl%3Den%26sa%3DG]
Ratarda formosana" Matsumura, 1921
Ratarda furvivestita" Hampson
Ratarda guttifera" Hering, 1925
Ratarda javanica" Roepke, 1937
Ratarda marmorata" Moore, 1879
Ratarda melanoxantha" Hering, 1925
Ratarda mora" Hering, 1925
Ratarda tertia" Strand, 1916
Ratarda monstrosa" Strand, 1916
Ratarida excellens" (Strand, 1917) (originally in "Shisa")
Sumatratarda diehlii" Kobes and Ronkay, 1990.
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