Melanella

Melanella
Melanella candida
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
(unranked):	clade Caenogastropoda clade Hypsogastropda Group Ptenoglossa
Superfamily:	Eulimoidea
Family:	Eulimidae
Genus:	Melanella Bowdich, 1822
Synonyms
Balcis Leach, 1847 Eulimaustra Laseron, 1955

Melanella is a genus of very small ectparasitic sea snails, marine gastropod mollusks or micromollusks in the family Eulimidae. ^[1]

Species

According to the World Register of Marine Species (WoRMS) the following species with accepted names are included within the genus Melanella ^[1]

• Melanella abida Dall, 1927
• Melanella acicula (Gould, 1849)
• Melanella aciculata (Pease, 1860)
• Melanella acuformis (G. Nevill & H. Nevill, 1875)
• Melanella aequatorialis (Thiele, 1925)
• Melanella aethiopica (Thiele, 1925)
• Melanella aethria (Melvill, 1918)
• Melanella alayoi Espinosa, Ortea & Magana, 2001
• Melanella alba (da Costa, 1778)
• Melanella alfredensis Bartsch, 1915
• Melanella amabilis (Brazier, 1877)
• Melanella amblytera (Verrill & Bush, 1900)
• Melanella ameliae Engl, 2007
• Melanella anachorea Dall, 1927
• Melanella angulata (A. Adams, 1861)
• Melanella antarctica (Strebel, 1908)
• Melanella arleyi Espinosa, Ortea & Magana, 2001
• Melanella atlantica (E. A. Smith, 1890)
• Melanella attenuata (Sowerby, 1866)
• Melanella atypha (Verrill & Bush, 1900)
• Melanella augur (Angas, 1865)
• Melanella aurata Hirase, 1920
• Melanella baldra Bartsch, 1917
• Melanella becki (Turton, 1932)
• Melanella bermudezi Pilsbry & Aguayo, 1933
• Melanella borealis (Bartsch, 1915)
• Melanella boscheineni Engl, 2004
• Melanella bovicornu (Pilsbry, 1905)
• Melanella brevis (Requien, 1848)
• Melanella breviuscula (Dunker, 1875)
• Melanella callistemma Dall, 1927
• Melanella catalinensis Bartsch, 1917
• Melanella chariessa (A. E. Verrill, 1884)
• Melanella charissa (Verrill, 1884)* : misspelling of Melanella chariessa (A. E. Verrill, 1884)
• Melanella chascanon (Watson, 1883)
• Melanella chaunax (Watson, 1883)
• Melanella chrysallida (A. Adams, 1861)
• Melanella chydaea (Watson, 1883)
• Melanella cifara Bartsch, 1915
• Melanella cinca Dall, 1927
• Melanella clavula (A. Adams, 1861)
• Melanella columbiana (Bartsch, 1917)
• Melanella compactilis (Monterosato, 1884)
• Melanella compsa (Verrill & Bush, 1900)
• Melanella conoidea (Kurtz & Stimpson, 1851)
• Melanella convexa (Smith, 1907)
• Melanella cosmia Bartsch, 1917
• Melanella costellata (Dautzenberg & Fischer H., 1897)
• Melanella cumingi (A. Adams, 1854)
• Melanella cuspidata (A. Adams, 1851)
• Melanella cylindrata (Watson, 1883)
• Melanella debilis (A. Adams, 1861)
• Melanella decagyra (Melvill & Standen, 1903)
• Melanella decipiens (Hedley, 1899)
• Melanella densicostata Bouchet & Warén, 1986
• Melanella doederleini (Brusina, 1886)
• Melanella draconis Bartsch, 1917
• Melanella dufresnii Bowdich, 1822
• Melanella eburnea (A. Adams, 1861)
• Melanella elenensis (Bartsch, 1924)
• Melanella elsa Thiele, 1930
• Melanella eulimoides (C. B. Adams, 1850)
• Melanella exasperata (Preston, 1905)
• Melanella exilis (Pease, 1863)
• Melanella exulata (Smith, 1915)
• Melanella falcata (Carpenter, 1865)
• Melanella fasciata (Watson, 1883)
• Melanella frielei (Jordan, 1895)
• Melanella fuscopunctata (E. A. Smith, 1890)
• Melanella gagei Bouchet & Warén, 1986
• Melanella glypta Bouchet & Warén, 1986
• Melanella gomphus (Watson, 1883)
• Melanella gracilis (C. B. Adams, 1850)
• Melanella gradata (Cotton & Godfrey, 1932)
• Melanella grippi (Bartsch, 1917)
• Melanella guenteri Engl, 2004
• Melanella guildingii (A. Adams, 1854)
• Melanella hadalis Knudsen
• Melanella halia Bartsch, 1917
• Melanella haloraphe (Dautzenberg & Fischer H., 1896)
• Melanella hastata (Sowerby, 1834)
• Melanella hebes (Watson, 1883)
• Melanella hemphilli Bartsch, 1917
• Melanella hians (Watson, 1883)
• Melanella hyalina (Watson, 1883)
• Melanella hypsela (Verrill & Bush, 1900)
• Melanella ima Pilsbry, 1918
• Melanella inconspicua (Watson, 1897)
• Melanella indica (Preston, 1905)
• Melanella insculpta Bouchet & Warén, 1986
• Melanella iotoides Bartsch, 1916
• Melanella ira Dall, 1927
• Melanella junieri Espinosa, Ortea & Magana, 2006
• Melanella kahoolawensis Pilsbry, 1917
• Melanella kanaka (Pilsbry, 1917)
• Melanella karolinae Engl, Winfried, 2006
• Melanella kawamurai (Kuroda & Habe, 1950)
• Melanella laseroni Hedley, 1916
• Melanella lastra Bartsch, 1917
• Melanella laurae (Friele, 1886)
• Melanella letsonae (Pilsbry, 1917)
• Melanella lineata (Monterosato, 1869)
• Melanella lubrica (Monterosato, 1890)
• Melanella lucida (Verrill, 1884)
• Melanella luminosa Marshall, 1997
• Melanella lunata Pilsbry, 1917
• Melanella lutea Rodrigez Garcia del Castillo, 2000
• Melanella macra Bartsch, 1917
• Melanella magnifica (Bartsch, 1915)
• Melanella maracuya Espinosa, Ortea & Magana, 2001
• Melanella martinii (A. Adams in Sowerby, 1854)
• Melanella martynjordani (Jordan, 1895)
• Melanella marviva Espinosa, Ortea, & Magaña, 2006
• Melanella medipacifica (Pilsbry, 1917)
• Melanella micans (Carpenter, 1865)
• Melanella microsculpta Bouchet & Warén, 1986
• Melanella mimus Pilsbry, 1918
• Melanella minor (Pallary, 1904)
• Melanella modesta Thiele, 1930
• Melanella montereyensis Bartsch, 1917
• Melanella muelleriae (Sturany, 1904)
• Melanella mundula (A. Adams, 1861-b)
• Melanella myriotrochi Bouchet & Warén, 1986
• Melanella nilae Engl, Winfried, 2006
• Melanella nisonida (Melvill, 1906)
• Melanella nutans (Mühlfeld, 1824)
• Melanella obtusa (Laseron, 1955)
• Melanella obtusoapicata Bouchet & Warén, 1986
• Melanella ochsneri (Bartsch, 1917)
• Melanella odontoidea (A. Adams, 1861)
• Melanella ogasawarana (Pilsbry, 1905)
• Melanella oleacea (Kurtz & Stimpson, 1851)
• Melanella olssoni (Bartsch, 1924)
• Melanella opaca (Sowerby, 1865)
• Melanella opalina (Monterosato MS, Marshall, 1901)
• Melanella ophiodon Dall, 1927
• Melanella orphanensis Clarke A.H., 1974
• Melanella orthophyes (Sturany, 1903)
• Melanella oxytata (Watson, 1883)
• Melanella pandata (A. Adams, 1861)
• Melanella parallela Dall, 1927
• Melanella peninsularis (Bartsch, 1917)
• Melanella penna Dall, 1927
• Melanella pernula (Monterosato, 1890)
• Melanella perplexa Oliver, 1915
• Melanella persimilis (Kuroda & Habe in Kuroda et al., 1971)
• Melanella perversa (Bush, 1909)
• Melanella petitiana (Brusina, 1869)
• Melanella pisinna Rehder, 1980
• Melanella planisutis Bouchet & Warén, 1986
• Melanella platyacme (Sykes, 1903)
• Melanella polita (Linnaeus, 1758)
• Melanella praecurta (Pallary, 1904)
• Melanella producta (Carpenter, 1863)
• Melanella pseudoglabra (Dautzenberg & Fischer H., 1912)
• Melanella psila (Watson, 1883)
• Melanella pyramidalis (A. Adams, 1854 in H. & A. Adams 1853-58)
• Melanella retexta (Carpenter, 1863)
• Melanella reticulata Bouchet & Warén, 1986
• Melanella retrorsa (Sowerby, 1865)
• Melanella richardi (Dautzenberg & Fischer H., 1896)
• Melanella robusta (A. Adams, 1861)
• Melanella roegerae (Cotton & Godfrey, 1932)
• Melanella rubrotincta (Jeffreys, 1867)
• Melanella salvadori Castellanos, Rolán & Bartolotta, 1987
• Melanella sankurieae Engl, 2004
• Melanella sansibarica (Thiele, 1925)
• Melanella sarissa (Watson, 1883)
• Melanella sarsi (Bush, 1909)
• Melanella seguenzai Di Geronimo & La Perna, 1999
• Melanella shibana (Yokoyama, 1927)
• Melanella siberutensis (Thiele, 1925)
• Melanella similior Bouchet & Warén, 1986
• Melanella sinuosa (Scacchi, 1836)
• Melanella solida (Sowerby, 1865)
• Melanella solidula (Adams & Reeve, 1850)
• Melanella soliduloides (Habe, 1951)
• Melanella solitaria (Smith, 1915)
• Melanella spina (Boettger, 1893)
• Melanella spinosa Oliver, 1915
• Melanella spiridioni (Dautzenberg & Fischer H., 1896)
• Melanella stalioi (Brusina, 1869)
• Melanella stamina Dall, 1927
• Melanella subantarctica (Strebel, 1908)
• Melanella subula (A. Adams, 1861)
• Melanella teinostoma (A. Adams, 1853)
• Melanella teramachii (Habe, 1952)
• Melanella thalia Bartsch, 1915
• Melanella tia (Bartsch, 1926)
• Melanella tortuosa (Adams & Reeve, 1850)
• Melanella townsendi Bartsch, 1917
• Melanella translucens (Monterosato, 1890)
• Melanella trunca (Watson, 1897)
• Melanella tumidula (Thiele, 1912)
• Melanella turbonilloides Bouchet & Warén, 1986
• Melanella turrita Bouchet & Warén, 1986
• Melanella vegrandis (Murdoch & Suter, 1906)
• Melanella versa Dall, 1927
• Melanella xiphidiopsis (Dautzenberg & Fischer, 1896)
• Melanella yamazii (Habe, 1952)
• Melanella zugnigae Espinosa, Ortea & Magana, 2001

Melanella candida, abapertural view

The Indo-Pacific Molluscan Database also includes the following species with names in current use ^[2]

• Melanella acanthyllis (Watson, 1886)
• Melanella acerrima (Watson, 1883)
• Melanella acuta (Sowerby, 1834)
• Melanella angulata (A. Adams, 1861-b)
• Melanella anomala (Laseron, 1955)
• Melanella australiensis (Thiele, 1930)
• Melanella bovicornu (Pilsbry, 1905)
• Melanella brunnimaculata (Kay, 1979)
• Melanella campyla (Watson, 1883)
• Melanella carneola (Gould, 1861 in 1859-61)
• Melanella clypeastericola (Habe, 1976-c)
• Melanella cunaeformis (May, 1915)
• Melanella curvata (A. Adams, 1861-b)
• Melanella dentaliopsis (A. Adams, 1861-b)
• Melanella flexa (A. Adams, 1861-b)
• Melanella flexuosa (A. Adams in H. & A. Adams, 1853-58)
• Melanella grandis (A. Adams, 1854 in H. & A. Adams, 1853-58)
• Melanella inflata (Tate & May, 1900)
• Melanella inflexa (Pease, 1868-b)
• Melanella labiosa (Sowerby II, 1834-b)
• Melanella lactea (A. Adams, 1854-i)
• Melanella latipes (Watson, 1883)
• Melanella legrandi (Beddome, 1882)
• Melanella mabutii Nomura & Hatai, 1935
• Melanella mamilla (May, 1915)
• Melanella maui (Del, 1952)
• Melanella modicella (Adams, 1854)
• Melanella montebelloensis (Iredale, 1914)
• Melanella montageuana (Iredale, 1914)
• Melanella musta (Yokoyama, 1928)
• Melanella nitens (Brazier, 1877)
• Melanella nivea (A. Adams, 1861-b)
• Melanella oblonga (Boettger)
• Melanella peronellicola (Kuroda & Habe, 1950-d)
• Melanella pinguicula (A. Adams, 1861-b)
• Melanella polygyra (Adams, 1854)
• Melanella reclinata (A. Adams, 1861-b)
• Melanella schoutanica (May, 1915)
• Melanella scitula (A. Adams, 1861-b)
• Melanella semitorta (A. Adams, 1861-b)
• Melanella stenostoma (A. Adams, 1861-b)
• Melanella stylata (A. Adams, 1861-b)
• Melanella tenisoni (Tryon, 1886)
• Melanella tryoni (Tate & May, 1900)
• Melanella valida (A. Adams, 1861-b)
• Melanella venusta (Pease, 1868-b)
• Melanella victoriae (Gatliff & Gabriel, 1914)

Species brought into synonymy 

• Melanella boscii (Payraudeau, 1826) : synonym of Melanella polita (Linnaeus, 1758)
• Melanella bryani (Pilsbry, 1917) : synonym of Melanella aurata Hirase, 1920
• Melanella candida (Marrat, 1880) : synonym of Melanella martinii (A. Adams in Sowerby, 1854)
• Melanella conoidalis (Sowerby, 1865) : synonym of Melanella cumingi (A. Adams, 1851)
• Melanella corrida Dall, 1927 : synonym of Melanella abida Dall, 1927
• Melanella crosseana (Brusina, 1886) : synonym of Eulima crossei (Brusina, 1886)
• Melanella engonia (Verrill & Bush, 1900)  : synonym of Eulima hypsela Verrill & Bush, 1900
• Melanella helena Thiele, 1930 : synonym of Melanella aciculata (Pease, 1861)
• Melanella indeflexa (A. Adams, 1861) : synonym of Melanella dufresnei (Bodwich, 1822)
• Melanella intermedia (Cantraine, 1835) : synonym of Melanella polita (Linnaeus, 1758)
• Melanella jamaicensis (C. B. Adams, 1845) : synonym of Melanella eburnea (Mühlfeld, 1824)
• Melanella jeffreysi (Tryon, 1886) : synonym of Melanella platyacme (Sykes, 1903)
• Melanella kuronamako (Habe, 1952)  : synonym of Melanella aciculata (Pease, 1861)
• Melanella kyurokusimensis Nomura & Hatai, 1960 : synonym of Melanella odontoidea (A. Adams, 1861)
• Melanella lampra Melvill, 1918 : synonym of Oceanida lampra (Melvill, 1918)
• Melanella luchuana (Pilsbry, 1901) : synonym of Melanella cuspidata (A. Adams, 1851)
• Melanella mayi (Tate, 1900) : synonym of Melanella augur (Angas, 1865)
• Melanella monterosatoi (Monterosato, 1890) : synonym of Melanella pyramidalis (Sowerby, 1866)
• Melanella murrayae (Cotton & Godfrey, 1932) : synonym of Melanella augur (Angas, 1865)
• Melanella orthopleura (Tate, 1898) : synonym of Melanella augur (Angas, 1865)
• Melanella planicincta (Cotton & Godfrey, 1932) : synonym of Melanella augur (Angas, 1865)
• Melanella proxima (Sowerby, 1866) : synonym of Melanella augur (Angas, 1865)
• Melanella solida (Sowerby, 1865) : synonym of Melanella platyacme (Sykes, 1903)
• Melanella solidula (Adams and Reeve, 1850) : synonym of Melanella lubrica (Monterosato, 1890)
• Melanella thaanumi Pilsbry, 1917 : synonym of Eulima major G.B. Sowerby, 1834
• Melanella vafra Pilsbry, 1917 : synonym of Parvioris saccata (Boettger, 1893)

1. ^ ^{a b} Gofas, S. (2011). Melanella Bowdich, 1822. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=137978 on 2011-05-20
2. ^ OBIS : Melanella

• Warén A. (1984) A generic revision of the family Eulimidae (Gastropoda, Prosobranchia). Journal of Molluscan Studies suppl. 13: 1-96. page(s): 54
• Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 180-213