Polystichum aculeatum

Taxobox
name = "Polystichum aculeatum"

---

image_width = 240px
regnum = Plantae
divisio = Pteridophyta
classis = Pteridopsida
ordo = Polypodiales
familia = Dryopteridaceae
genus = "Polystichum"
species = "P. aculeatum"
binomial = "Polystichum aculeatum"
binomial_authority = (L.) Roth

"Polystichum aculeatum" (Hard Shield Fern) is an evergreen fern native to Europe. It is most abundant in upland regions of the British Isles and western France, where it benefits from the combination of mild winters and moist summers, but also occurs more locally across most of Europe except northern Scandinavia, northern Russia; in the Mediterranean region it is confined to high altitudes. It grows on steep slopes in woodlands.

The glossy dark green fronds are 30-90 cm long, usually drooping downslope, with typically 3-8 fronds on a mature plant. The fronds are stiff and hard-textured, bipinnate (single-pinnate on small, young plants), with the pinnae opposite on the stalk. Each pinna is 3-11 cm long, with a large upward-pointing pinnule at the base, and the other pinnules decreasing in size toward the pinna tip; the pinnules have bristly tips. Individual fronds live for 1.5-2.5 years and remain attached to the rhizome after withering. The round sori occupy two rows on either side of the midrib of each pinnule and are covered by a centrally-attached, umbrella-like indusium with fringed edges. They produce light yellow spores.

It is occasionally grown as an ornamental plant in gardens.

References

* [http://rbg-web2.rbge.org.uk/cgi-bin/nph-readbtree.pl/feout?FAMILY_XREF=&GENUS_XREF=Polystichum+&SPECIES_XREF=aculeatum&TAXON_NAME_XREF=&RANK= Flora Europaea: "Polystichum aculeatum"]
*Hyde, H. A., Wade, A. E., & Harrison, S. G. (1978). "Welsh Ferns". National Museum of Wales.
*Swan, G. A. (1993). "Flora of Northumberland". Natural History Society of Northumbria.