Heptasteornis

Taxobox|
name = "Heptasteornis"
fossil_range = Late Cretaceous (Maastrichtian)
regnum = Animalia
phylum = Chordata
classis = Sauropsida
superordo = Dinosauria
ordo = Saurischia
subordo = Theropoda
unranked_familia = Coelurosauria
familia = Alvarezsauridae
genus = "Heptasteornis"
genus_authority = Harrison & Walker, 1975
species = "H. andrewsi"
binomial = "Heptasteornis andrewsi"
binomial_authority = Harrison & Walker, 1975

"Heptasteornis" is the name given to a dubious genus of small dinosaur from the Late Cretaceous. The type (and only known) species is "Heptasteornis andrewsi", described as a presumed gigantic prehistoric owl in 1975. It was previously included in "Elopteryx nopcsai" and indeed the holotypes of both were believed to be from the same individual as they were discovered, and initially were assigned the same specimen number. This appears to be in error however (see below).Harrison & Walker (1975)]

The material was discovered in Romania, in the late Maastrichtian Sânpetru Formation (Rognacian faunal stage, deposited c. 68-65.5 million years ago) of the Haţeg Basin in Transylvania [Mortimer (2004)] . The scientific name means "C.W. Andrews' Transylvanian bird", after the discoverer of the first specimen [Andrews (1913)] and Ancient Greek "(h)epta" (επτά) "seven" + "asty" (άστυ) "city" + "ornis" (όρνις) "bird"; the Latin "septum urbium" or the German "Siebenbürgen" - meaning "seven cities" or "seven castles" - were common names for the Transylvanian region throughout the centuries. The material was originally limited to a mere two broken distal tibiotarsi, BMNH A4359 (the holotype) and A1528. The taxonomic status and systematic placement of these bones was much disputed and they were often considered junior synonyms of "Bradycneme" or "Elopteryx". Given the fragmentary nature of the fossils, little could be resolved and "Heptasteornis" was (and still is) considered a "nomen dubium" by many. [Paul (1988), Weishampel "et al." (1991), Le Loeuff "et al." (1992), Csiki & Grigorescu (1998)]

However, more recently the bones were reassessed as an alvarezsaurid, the first to be known from Europe. Not only has this theory, originally proposed in 1988, withstood further scrutiny, but it appears as if "H. andrewsi" quite clearly belongs to the subfamily Mononykinae. "Bradycneme" and "Elopteryx" on the other hand seem to be more advanced maniraptoran theropods. The results of the restudy of BMNH A4359 also suggest that a distal piece of femur, specimen FGGUB R.1957, probably can also be assigned to "Heptasteornis". [Mortimer (2004), Naish & Dyke (2004)]

Thus, quite ironically, of those three enigmatic Romanian theropods the one most explicity named a "bird" - as per its scientific name - is almost certainly the one most distantly related to birds.

Footnotes

References

* (1913): On some bird remains from the Upper Cretaceous of Transylvania. "Geological Magazine" 5: 193-196.
* (1998): Small theropods from the Late Cretaceous of the Hateg Basin (western Romania) - an unexpected diversity at the top of the food chain. "Oryctos" 1: 87-104.
* (1975): The Bradycnemidae, a new family of owls from the Upper Cretaceous of Romania. "Palaeontology" 18(3): 563-570. [http://palaeontology.palass-pubs.org/pdf/Vol%2018/Pages%20563-570.pdf PDF fulltext]
* (1992): The first record of dromaeosaurid dinosaurs (Saurischia, Theropoda) in the Maastrichtian of southern Europe: palaeobiogeographical implications. "Bulletin de la Société géologique de la France" 163(3): 337-343.
* (2004): The Theropod Database: [http://home.comcast.net/~eoraptor/Phylogeny%20of%20Taxa.html Phylogeny of taxa] . Retrieved 2008-AUG-14.
* (2004): "Heptasteornis" was no ornithomimid, troodontid, dromaeosaurid or owl: the first alvarezsaurid (Dinosauria: Theropoda) from Europe. "Neues Jahrbuch für Geologie und Paläontologie Monatshefte" 7: 385-401.
* (1988): "Predatory Dinosaurs of the World." New York, Simon & Schuster. ISBN 0-671-61946-2
* (1991): The dinosaurs of Transylvania. "National Geographic Research and Exploration" 7(2): 196-215. [http://bio.fsu.edu/~amarquez/Evolutionary%20Morphology%20fall%202004/Weishampel/1096-%20Weishampel%20et%20al%201991%20-%20Transylvania%20dinosaurs.pdf PDF fulltext]

External links

* http://www.dinoruss.com/de_4/5a8b459.htm
* http://www.users.qwest.net/~jstweet1/coelurosauria.htm